ana Zuece. MAP 19b 
ingeana Zucc. in Endl., Synops. Conif. 
166. 1847. 
rk pine grows in northern China from northern 
nd Hupeh north to Shansi and Hopeh. Notable 
its peculiar Platanus-like bark, this pine is com- 
rare throughout its range. 
ublished—Cheng 1939 (Szechuan); H. F. Chow 1934 
peh) ; Hers 1922 (Honan); Hu 1935 (Honan); H. Smith 
5 (Shansi); Sowerby 1937; ‘Tang 1931 (Shansi) ; Teng 1947 
su); Wilson 1907 (Hupeh). 
Unpublished—Wang Chi-Wu 1964 (Shensi). 
SUBSECT. BALFOURIANAE Shaw 
Pinus balfouriana Grev. & Balf. MAP 20 
foxtail pine (see also Map 21) 
Pinus balfouriana Grev. & Balf. in A. Murr., Bot. 
Exped. Oreg. | Rpt. No. 8] No. 618, t. 1853. 
Foxtail pine is confined to two high-elevation areas about 
300 miles apart: the southern Sierra Nevada, Calif., and the 
higher mountains of northwestern California from the South 
Yolla Bolly Mountains north to the Marble Mountains. The 
stands on the east slope of the Sierra Nevada are about 20 
miles from stands of Pinus aristata in the Inyo Mountains, 
only the width of the arid Inyo Valley separating the two 
species at this point. A northern outlier of the Sierra Nevada 
distribution reported by Sudworth (1908, p. 41) on the 
divide between the South Fork of the San Joaquin River and 
the North Fork of the Kings River has never been verified, 
and is not included in the distribution shown on the map. 
Additional sources: 
Published—Howell 1951. 
File reports—“Survey of foxtail pine in northern Cali- 
fornia” by T. H. Harris and Benton Howard, August 28, 1956. 
Forest Service, Region 5, San Francisco, Calif.; “Blister rust 
occurrence survey in the Marble Mountains, Klamath Na- 
tional Forest’”’ by Benton Howard, October 4, 1956. Forest 
Service, Region 5, San Francisco, Calif. 
Unpublished—D. V. Hemphill 1964 (northern Calif.) . 
MAP 21 
(see also Map 20) 
Pinus aristata Engelm. in Parry & Engelm., Amer. 
Jour. Sci. and Arts, Ser. 2, 34: 331. 1862. 
Bristlecone pine ranges from eastern California through 
Nevada, Utah, and northern Arizona to Colorado and north- 
ern New Mexico. It grows at high elevations and frequently 
extends to timberline. In many places it is very localized, 
but in others, such as the White Mountains of eastern Cali- 
fornia, it forms extensive stands. It is probably more widely 
distributed in the poorly known mountain ranges of Nevada 
than the map indicates. 
Additional sources: 
Unpublished—W. P. Cottam 1964 (Utah); Margaret M. 
Douglass 1962 (Colo.); C. W. Ferguson 1962 (Calif.); R. D. 
Wright 1962 (Calif.). 
Pinus aristata Engelm. 
bristlecone pine 
10 
SUBGEN. PINUS 
SECT. TERNATAE Loud. 
SUBSECT. LEIOPHYLLAE Loud. 
Pinus leiophylla Schiede & Deppe MAP 22 
Chihuahua pine 
Pinus letophylla Schiede & Deppe in Schlecht. & 
Cham., Linnaea 6: 354. 1831. 
Chihuahua pine ranges from southern and central Mexico 
through western Mexico to southwestern New Mexico and 
southern and central Arizona. Included here is Pinus leio- 
phylla var. chihuahuana (Engelm.) Shaw, sometimes consid- 
ered a separate species, P. chihuahuana Engelm. P. leiophylla 
is unique among American pines in having cones that require 
three seasons to mature. Its northernmost known occurrences 
are just south of the Mogollon Rim in central Arizona (7 
miles west of Pinedale—Sudworth 1927; about 25 miles east 
of Payson—M. B. Applequist, personal communication, 1964). 
Additional sources: 
Published—Goldman 1951; Guzman and Vela 
(Zac.) ; J. IT. Marshall 1957 (Son., Chih.) ; Shaw 1909. 
Thesis—James H. Maysilles 1959. Floral relationships 
of the pine forest of western Durango, Mexico. Ph.D. thesis, 
Univ. Michigan, 165 pp., illus. 
Unpublished—M. B. Applequist 1964 (Ariz.); B. Hall- 
berg 1964 (Oax.); E. Larsen 1962; N. Sanchez Mejorada to 
L. W. Bryan 1963 (Mich.); O. Zarzosa L. 1964 (Dgo.). 
1960 
Pinus lumholtzii Robins. & Fern. MAP 23 
Lumholtz pine 
Pinus lumholtzii Robins. & Fern., Amer. Acad. 
Proc. 30: 122. 1895. 
This species grows at relatively low elevations in north- 
western Mexico, from Sonora and Chihuahua south to Jalisco. 
It is notable for its pendent but stiff needles. 
Additional sources: 
Published—Gentry 1942, 1946 (Sin.); Guzman and Vela 
1960 (Zac.); Shaw 1909 (Nay.). 
Thesis—James H. Maysilles 1959. Floral relationships of 
the pine forests of western Durango, Mexico. Ph.D. thesis, 
Univ. Michigan, 165 pp., illus. 
Unpublished—C. E. Blanco 1964 (Dgo.) ; E. Larsen 1962; 
O. Zarzosa L. 1964 (Dgo.). 
SUBSECT. CANARIENSES Loud. 
Pinus canariensis C. Smith MAP 24b 
Canary Island pine 
Pinus canariensis C. Smith in Buch, Phys. Beschr. 
Canar. Ins. 159. 1825. 
Canary Island pine is confined to the central and western 
Canary Islands, off the coast of northwestern Africa. Its only 
close relative, Pinus roxburghii, grows in the Himalaya 
Mountains more than 5,000 miles away. 
Sources: 
Published—Ceballos and Ortuno 1951. 
Unpublished—L. Ceballos to N. T. Mirov (n.d.) (Gran 
Canaria). 
