SPECIES DISTRIBUTION, GENUS PINUS L. 
SUBGEN. DUCAMPOPINUS (A. Cheval.) 
de Ferré 
SECT. DUCAMPOPINUS 
SUBSECT. KREMPFIANI Little & Critehfield’ 
Pinus krempfii Lecomte MAP 2a 
Pinus krempfii Lecomte, Paris Mus. Natl. d’Hist. 
Nat. Bull 27: 191, fig. 1921: 
This unique species—the only pine with flattened needles 
—is known only from the high mountains of southern Viet- 
nam. 
Sources: 
Published—Bui 1962; Rollet 1955. 
Unpublished—M. Schmid to N. T. Mirov 1961. 
SUBGEN. STROBUS Lemm. 
SECT. STROBUS 
SUBSECT. CEMBRAE Loud. 
Pinus koraiensis Sieb. & Zuce. MAP 2b 
Korean pine (see also Map 11) 
Pinus koraiensis Sieb. & Zucc., Fl. Jap. 2: 28, 
t. 116, fig. 5-6. 1844; exclud. fig. 1-4. 
Korean pine ranges through Korea and eastern Man- 
churia into southeastern Siberia, with outliers on the Japanese 
islands of Honshu and Shikoku. 
Sources: 
Published—Hayashi 1952 (Japan); Solovev 1958 
(U.S.S.R.); Uyeki 1926 (Korea); Wu 1956 (Manchuria). 
Pinus pumila Regel MAP 3 
Japanese stone pine (see also Map 4) 
Pinus pumila Regel, Index Sem. Hort. Petrop. 
1858: 23. 1859. 
This pine ranges widely through northwestern Asia, ex- 
tending north almost to the Arctic Ocean, east to the Bering 
Sea, west to northern Mongolia and Lake Baikal, and south 
to Korea and central Honshu, Japan. In the southern part 
of its range this shrubby, thicket-forming species is restricted 
to high elevations. Its presence on the Commander Islands 
east of Kamchatka is debatable; Tikhomirov (1949) shows it 
on one of the islands (Mednyy), but Tatewaki (1958) states 
that it is not present on either island. 
' Pinus subgen. Ducampopinus (A. Cheval. ) de Ferre, subgen. nov. 
Pinus sous-genre Ducampopinus (A. Cheval.) de Ferré, Paris Acad. Sci. 
Compt. Rend. 236: 228. 1953 (not validly published under [CBN Art. 33 
because reference to basionym lacked original publication with page) . 
Ducampopinus A. Cheval., Rev. Bot. Appl. d’Agr. Trop, 24: 30. 1944. 
Holotype species: Pinus krempfit Lecomte, Paris Mus. Natl, Hist. Bul. 27: 
LOE tiga O21 
5 Pinus subgen. Ducampopinus sec, Ducampopinus subsect. Kremp/fiani 
Little & Critchfield, subsect. nov. Characteribus subgeneris Pinus subgen. 
Ducampopinus (A, Cheval.) de Ferré. Folio cum 1 fasciculo vasculare, 2 
In the past this species has generally been considered a 
variant of the Pinus cembra—P. sibirica group of eastern Si- 
beria and Europe, but recent investigators consider it more 
closely allied to the Japanese P. parviflora (de Ferré 1960; 
Malyshev 1960) . 
Sources: 
Published—Hultén 1926; Imanishi 1950 (Manchuria) ; 
Kung 1934 (Manchuria); Malyshev 1960 (Siberia); Takahasi 
1944 (Manchuria); Tatewaki 1958 (Kurile Islands); Tik- 
homirov 1949; Utkin 1961 (Siberia); Uyeki 1926 (Korea) ; 
Wu 1956 (Manchuria) . 
Pinus sibirica Du Tour MAP 4 
Siberian stone pine 
Pinus sibirica Du Tour, Nouv. Dict. Hist. Nat. 18: 18, 1803. 
Siberian stone pine ranges from the Ural Mountains 
through western and central Siberia to northern Mongolia, 
with outliers on the Kola Peninsula of northern Russia. This 
species is the Siberian counterpart of the central European 
Pinus cembra. Its range overlaps that of P. pumila in the 
region of Lake Baikal, Siberia. P. stbirica sometimes assumes 
a shrubby growth form like that of P. pumila, and the two 
are often confused (Malyshev 1960). 
Sources: 
Published—Hultén 1950 (Kola Peninsula); Lavrenko and 
Soczava 1956; Nekrasova 1949 (Kola Peninsula) . 
Pinus cembra L. MAP 4 
Swiss stone pine 
Pinus cembra L., Sp. Pl. 1000. 1753. 
Swiss stone pine grows at high elevations in the Alps and 
the Carpathian Mountains. It is closely related to Primus st- 
birica, from which it is separated by a 1,500-mile gap. 
Sources: 
Published—Fekete and Blattny 1913-1914; Generali 1937 
(Italy); Marchesoni 1959 (Italy); Ozenda 1961 (France) ; 
Rubner 1953, p. 364; Scharfetter 1938; Schmid 1949 (Switzer- 
land) . 
Unpublished—Forestry Research Institute, Rumania, 
1964; M. Vidakovié 1964 (Yugoslavia). 
Pinus albicaulis Engelm. MAP 5 
whitebark pine 
Pinus albicaulis Engelm., Acad. Sci. St. Louis Trans. 
2: 209. 1863. 
Whitebark pine occurs at high elevations from central 
British Columbia south to the southern Sierra Nevada olf 
in fasciculo, anguste lanceolata, valde complanata (15-4 mm. late 
minute serrulata vel integra, stomatibus ventralibus atque raro dorsalibus 
ductis resiniferis subexternis, et vagina mox decidua. Bases bractearum 
non decurrentes. Strobilus junioris solitarius, sine aculeis. Strobili ovoid 
symmetricales, aperti postquam maturi. Squamae carinatae, apophysi 
crassa pyramidata et umbone dorsali. Semina cum ala longa articulata 
Lignum modice durum, leviter resinosum, annulis incrementi, sinc 
tracheidiis radiis, alburno albo luteolo, ligno interiore col monis. A 
subgeneribus aliis differt follis complanatis atque absen tracheidiarum 
radiis. Holotypus: Pinus kremp/it Lecomte, Paris Mus. Nz Hist. Nat 
Bul. 27: 191, fig 1921. 
or 
