56 Richard Franz, Judy Bauer and Tom Morris 



there were solution channels in the limestone large enough to accommodate 

 them. Unfortunately, there are no records of these species from intermediate 

 areas, although Means (1977) noted a cave along the Apalachicola 

 River that should be sampled. No troglobites were located during pre- 

 liminary surveys of caves in Dothan County, Alabama (John E. Cooper, 

 personal communication, Raleigh, North Carolina). Slight or no morpho- 

 logical differences have been noted between the populations of Cambarus 

 cryptodytes and Haideotriton wallacei from the two areas (Pylka and 

 Warren 1958, Means 1977, Hobbs 1981), which further supports the 

 continuous population hypothesis. 



WooDviLLE Fauna 



Species List — Remasellus parvus, **Crangonyx hobbsi, **Crangonyx 

 grandimanus, *Procambarus horsti, *P. orcinus.The Woodville Fauna 

 is associated with the Ocala Group limestones (Floridan aquifer) in 

 the eroded portions of the Tallahassee Hills and the Woodville Karst 

 Plain, along and below the Cody Scarp, respectively. It is bounded on 

 the west by the Apalachicola Coastal Lowlands (see Hendry and Sproul 

 1966). The eastern limits remain to be defined, although none of the 

 fauna is known to occur east of the Aucilla drainage. Hendry and 

 Sproul (1966) and Yon (1966) show a more or less continuous limestone 

 shelf below the Cody Scarp (Woodville Karst Plain) across southern 

 Leon and Jefferson counties. Lane (1986:32) describes this area as a 

 "flat to gently undulating surface of sand that overlies carbonate rock. 

 The carbonates, which lie at shallow depths of 30 feet or less, have 

 undergone extensive solution by groundwater. This plain exhibits karst 

 features that are still evolving, for example: many old, well developed 

 sinkholes that are either permanently or intermittently flooded (Big 

 Dismal Sink), disappearing streams and natural bridges (Natural Bridge), 

 Wakulla Springs, and new sinkholes reported periodically." 



The fauna appears to follow the riverine karsts associated with 

 the Wakulla-St. Marks rivers and the Wacissa River. In this way, the 

 endemic crayfishes parallel the distribution of the closely-related Pro- 

 cambarus pallidus that tracks the riverine karsts of the upper Suwannee 

 River and its tributaries. P. orcinus may be restricted to the Wakulla 

 drainage, whereas P. horsti may be more closely-tied to the St. Marks 

 and Wacissa drainages. The ecological relationships between them 

 remain unclear, particularly in lieu of an account that both crayfishes 

 co-exist in the Wakulla Springs Cave System (Morris 1989). More 

 collecting is necessary in caves of the Woodville Karst Plain to deter- 

 mine the actual geographic extent of this fauna and its ecological 

 specializations. 



