The Chonaphini 115 



stituted tribe Chonaphini. The chonaphine taxa therefore relate through 

 differentially shared aspects of all the principal features; two forms 

 that do not share one attribute share others, or relate through a third 

 form with which they share still other features. 



Separate taxonomic status for this assemblage was first proposed 

 by Verhoeff (1941), who established the Chonaphinae as a subfamily 

 in the "Leptodesmidae." Hoffman (1979) reduced the taxon to tribal 

 status and listed Chonaphe Cook, Montaphe Chamberlin, Semionellus 

 Chamberlin, and Metaxycheir Buckett and Gardner as component genera. 

 He and I (Shelley 1990) placed Tubaphe Causey in the Harpaphini, an 

 understandable misassignment because one gonopod of the holotype 

 of T. levii Causey is lost, the other is broken, and the descriptive 

 accounts and illustrations (Causey 1954^) do not begin to portray the 

 taxonomically important features. I (Shelley 1990) proposed Metaxycheir 

 pacifica for a chonaphine on Vancouver Island, British Columbia, 

 that I subsequently (Shelley 1993^) had to place in synonymy under 

 T. levii after recollecting the latter in the Olympic Mountains. Despite 

 the unfortunate proposal of a synonym, my description and illustrations 

 (Shelley 1990) make T. levii the only chonaphine taxon that has been 

 characterized in accordance with modern standards. Most were proposed 

 with terse accounts that merely validated the names, and one new 

 genus and six new species await description. The purpose of this 

 contribution, therefore, is to present modern descriptions and illustrations 

 for all chonaphine taxa except T. levii, where such is provided in the 

 account of M. pacifica (Shelley 1990). Acronyms of sources of preserved 

 study material are as follows: 



AMNH — American Museum of Natural History, New York, 



New York. 

 BYU — Monte L. Bean Life Science Museum, Brigham Young 



University, Provo, Utah. 

 CAS — California Academy of Science, San Francisco. 

 CMN — Canadian Museum of Nature, Ottawa, Ontario. 

 EIL — Zoology Department, Eastern Illinois University, Charleston. 

 FMNH — Field Museum of Natural History, Chicago, Illinois. 

 FSCA — Florida State Collection of Arthropods, Gainesville. 

 MCZ — Museum of Comparative Zoology, Harvard University, 



Cambridge, Massachusetts. 

 MPM — Milwaukee Public Museum, Milwaukee, Wisconsin. 

 NCSM — North Carolina State Museum of Natural Sciences, Raleigh. 

 NMNH — National Museum of Natural History, Smithsonian 



Institution, Washington, D. C. 



