6 
The intramatrical hypostroma with superficial ascomata is typical of the Polystomel- 
laceae ; the radial arrangement of the locul which are attached at several points, the 
paraphysate asci, and two-celled brown spores are characteristic of the genus Schneepia, 
to which this fungus must therefore be referred. 
2. Cycloschizon brachylaenae (Rehm), P. Henn. 
Engl. Bot. Jahrb. XXXIII, p. 39 (1902); Syll. Fung. XVII, p. 896. 
Syn. Schneepia brachylaenae Rehm, Hedwigia 1901, p. 173. 
On living leaves Brachylaena mernfolia, Bains Kloof, near Wellington, C.P., 12.11.10, 
Doidge [981]. 
On Brachylaena dentata, Port Elizabeth, West [761]; Van Stadens Pass, 9.3.16, 
Bottomley [9562]; Van Stadens Pass, 17.5.18, Pole Evans [11445]. 
On Brachylaena discolor, Lemana, Zoutpansberg District, 14.8.11, Doidge [1789] ; 
Duikerfontein, Natal, 10.10.11, Moon [1887]; Port Shepstone, Natal, 15.10.12, Pole 
Evans [5605]; Umgeni, near Durban, 15.11.16, V. d. Byl [10088]. 
On Brachylaena elliptica, Kast London, 24.11.17, Doidge [10912]. 
On Brachylaena sp., Rikath, Portuguese East Africa, September, 1918, Junod [11724]. 
Sydow [Ann. Myc. XIII (1915), p. 207], described this fungus in detail from the type 
specimen (Exsicc. Rabh. P., F. Eur. et extra-europ., 4264), which agrees in every respect 
with the material in the National Herbarium. ‘The original is evidently not quite mature, 
and in this connection Theissen and Sydow writes as follows :—' Die Art unterscheidet 
sich generisch nur durch die farblosen Sporen von Dielsiella ; da die Asken des Originals 
noch nicht ganz ausgereift sind, wire es nach diesem Exemplar nicht ausgeschlossen, 
dass die Sporen sich spiter braiinten; dan fiele die Gattung Cycloschizon mit Dvelsiella 
(1903) zusammen und letztere miisste gestrichen werden. Zu demselben Ergebnis kam 
V. Hohnel (Fragm. n. 634 und 635). . . . Das oben erwihnte Exemplar auf B. dentata 
scheint jedoch véllig ausgereift zu sein. Wir fandan hier schon ausserhalb der Schlauche 
zahlreiche, stets véllig hyaline Sporen, Wahrscheinlich stellt Cycloschizon daher eine gute 
Gattung dar.” 
After examining the abundant material detailed above, it is evident that the spores 
of Cycloschizon brachylaenae become fuscous, and later brown, at maturity. The genus 
Dielsiella is therefore identical with Cycloschizon and the species Dielsiella pritzelu P. Henn., 
and D. alyxiae (Mass) Th. and Syd. become Cycloschizon pritzelu (P. Henn.) and C. alyxie 
(Mass), respectively. 
An apparently undescribed species on Catha edulis also belongs to the genus Cyclo- 
schizon, and is described below. 
3. Cycloschizon fimbriatum n. sp. 
On living leaves of Catha edulis, Letaba Drift, Zoutpansberg District, 6.8.11, Doidge 
[1798] and [1833]; Rikath, Portuguese East Africa, 31.5.18, Junod [11682]; Louis 
Trichardt, 8.4.19, Putterill [11835]. 
Stromata amphigenous, scattered, circular, 1-5 to 2 mm. diam., surface dull black, 
crustaceous, with an irregular annular ridge marking the position of the loculi; attached 
to the leaf by a massive central foot, which is black and pseudo-cellular, arising from a 
dense hypostroma. The hypostroma consists of closely interwoven and packed hyphae, 
which completely destroy the palisade cells or the mesophyll cells in the immediate 
vicinity of the foot and penetrate right through the thickness of the leaf in the inter- 
cellular spaces. The increasing pressure of the hypostroma ruptures the epidermis, and 
at this point the foot is formed. | 
Except for the central foot, the stroma lies free on the leaf surface. The outer 
membrane is radial in structure, rather loosely interwoven, and composed of hyphae 
3-5-4 thick. At the periphery it spreads out into a short fringe of brown, radiating 
hyphae about 3-5 thick; these are undulating, branched, and sometimes more or less 
