48 



Fred C. Rohde, Mary L. Moser and Rudolf G. Arndt 



All three rivers are generally bordered by forest, although some land is 

 pasture. River substrate ranges from bedrock to boulders to cobble to silt. 



Fig. 4. French Broad and Nolichucky river systems with sites (dots) we sampled 

 from July 1991 to April 1995. Some dots overlap. 











TENNESSEE 



/ 



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/ 









Nolichucky River 



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N^^ 





i "-vt L. 







y 



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/**" 







French Broad River -*>Jfc=-; 









Cane Rfver jj. 





\ North Toerfiiver 



vt\ 



'w J 











/'jm 















-^ — ^- r ) L 













A South Toe River 



1 















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1 



1 1 



NORTH CAROLINA 









\f 







16 32 

 KILOMETERS 



METHODS AND MATERIALS 



We sampled 99 sites in 105 collections: 32 sites in the lower reach of 

 the French Broad River and some of its tributaries from May 1994 to May 1995, 

 and at 2 sites in the middle reach of the river in September 1993 (Fig. 4); 39 sites 

 in the Nolichucky River system between July 1991 and November 1994 (Fig. 4); 

 and 15 sites in the Dan River and in 2 of its tributaries 25 times from July 1992 

 to May 1995 (Fig. 3). We sampled most sites only once, although four sites each 

 in the French Broad and Nolichucky rivers were each sampled twice. We also 

 sampled 1 1 sites in the Virginia portion of the upper Dan River from November 

 1993 to October 1994 (Fig. 3). All site locations and dates are available from the 

 senior author. In addition, we include data from 146 collections made by others, 

 other personal communications, and from the literature. 



We sampled primarily with a backpack electroshocker and seine, using 

 the technique described by Jenkins and Burkhead (1975). Each site was elec- 

 trofished for 45- 190 minutes, i.e., until we believed that sampling had been com- 



