108 Troy A. Ladine and Angela Ladine 



to detect observable differences. Additionally, the behavior of the species being 

 studied must be addressed in selecting the size and number of scales to be 

 assessed. For example, species with large ranges will require large scales to 

 account for greater movement of individuals of these species. 



ACKNOWLEDGEMENTS - We thank Michael L. Kennedy for the use of the 

 Sherman traps and the Edward J. Meeman Biological Station for the use of facil- 

 ities. We thank the reviewers for their constructive comments on a previous ver- 

 sion of this manuscript. 



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