42 Jan S. Doughman 



than ta 5 . Hypopygium with a rectangular sternapodeme. Sternite IX 

 with a small tubercle protruding over base of each gonocoxite; anal 

 point (AnP) absent or a small, awl-like protrusion of tergite IX which is 

 not supported by oblique tergal apodemes. Pars ventralis small. Median 

 volsella (MVo) clavate and setigerous; superior volsella (SVo) a strongly 

 sclerotized and curving band (Figs. 3, 12). Gonocoxite tapering, nearly 

 parallel sided, with longish setae scattered on the lower two-thirds; 

 proximal half of inferior volsella (IVo) with a pile of microtrichia and 

 distal half with short, perpendicular setae and ending slightly free of 

 gonocoxite; and a second medial 'lobe' covered with short setae extend- 

 ing the length of the middle third of the gonocoxite. Gonostylus tubu- 

 lar, pubescent and with a few setae distally, terminally with thick chitin 

 surrounding its face below the perpendicular macroseta, and two strong, 

 perpendicular setae stand just proximal to macroseta (Figs. 3, 7). 



Female. — Antenna with six flagellomeres (first two somewhat 

 fused). Alula of wing with setae and anal lobe not rounded. Tergite IX 

 apparently large, segment X without setae, lateral parts darkly sclerot- 

 ized, postgenital plate apparently well developed, cerci hexagonal with 

 subequal sides, and three oval seminal capsules. (Serra-Tosio 1971:Fig. 

 124; Saether 1977, based on S. zavreli). 



Pupa. — Length 5-7 mm. Frontal apotome without cephalic tuber- 

 cles, but long frontal seta arising from each rugose spot. Thoracic horn 

 absent. Middle precorneal seta longer and thicker than the two, sub- 

 equal, flanking setae. Thoracic, antennal and wing sheaths smooth. Leg 

 sheath of Pending in a Z-curve. Tergite I (Tl) with only a polygonal 

 pattern and only L 2 and L 4 present. Tergites II-VII with an anterior 

 band of color only (apophyse) followed by some group shagreen border- 

 ing polygons; then spinulae dispersing evenly and ending in rows of 

 small, triangular spines on the posterior margin; finally rows of strong, 

 anteriorly pointing spines on conjunctiva II /III - VII / VIII (Fig. 4). L 2 

 dorsal. L |3 evenly spaced but moving toward lateroposterior on each 

 successive segment. L 3 and L 4 in close proximity. L-setae usually api- 

 cally forked, up to eleven branches on L 2 on VII and VIII. D-setae 

 small, dispersed, and usually simple. MD1 somewhat transverse and 

 MD2 longitudinal. Shagreen on sternites similar to that of tergites. 

 Segment IX with a fringe of very short setae on lateral border; each lobe 

 having a terminal, tooth-like ventral tubercle near the last of the three 

 straight macrosetae. Gonopodal sacs straight and not extending past 

 caudal margin. 



Larva (4th stadium). — Head quadrate, rather thinly sclerotized, 

 and color uniform luteous or darker. Mentum with a yellow or smoky, 

 subequally trisected ventromentum (median area) that is 6-7X the width 

 of the much darker first dorsomental (lateral) tooth; six to seven 



