44 Jan S. Doughman 



Larvae can be found in crude cases in pools or in reaches with laminar 

 flow. Their guts contain predominantly diatoms. Flight time is from 

 March to June when water reaches 10-15°C. This ecological summary 

 was taken from Serra-Tosio (1971), Johannsen (1937), Pagast (1947), 

 Ferrarese and Rosaro (1981) and conversations with U.S. Geological 

 Survey personnel and with David Lenat of the North Carolina Depart- 

 ment of Natural Resources and Community Development. 



KEYS TO MALES, PUPAE AND LARVAE OF SYMPOTTHASTIA 



MALES 



1. AnP awl-like, bare, 60-90 um long. Capitellum dusky. Palpal 3d segment 



without a keel of dark setae {fulva-gp.) 2 



AnP absent or conical, hairy and one or two spines apically (Figs. 9-1 1). 

 Capitellum clear. Palpal 3d segment expanded bearing a keel of dark 

 setae (zavreli-gp.) 3 



2. AR 2.4-2.6; A )3 800 um long. LR 73, 49, 53; P 3 with ta 4 110 um and 

 shorter than ta Palpal length 500 um; palp to face about 85. Clypeal 

 setae about 15. SVo straight distally (Fig. 13). Body and wing about 3 mm 



S.fulva (Jon.) 



AR 2.7-2.9; A ]3 995 um long. LR 88, 54, 52; P 3 with ta 4 170 um and 

 longer than ta y Papal length 850 um; palp to face 87-98. Clypeals 12-15. 

 SVo sigmoid (Fig. 3). Body 5-6 mm; wing 3.6 mm . . . . S. diastena (Subl.) 



3. AnP absent or minuscule and hairy. Clypeal setae absent. LR 81, 49, 59. 

 AR 2.4-3.0; A |3 1150 um long. Palp to face 67-72. VR 85-92. SVo near 

 Fig. 12, but lacking spinulae. Body 4-6 mm; wing 3-4 mm . . .S. zavreli Pag. 

 AnP conical, hairy, and stong spine(s) apically. Clypeals present. LR 75, 

 52, 58 or 64. VR 87-90 4 



4. AnP about 35 um long with one spine apically (Figs. 7, 9, 10). Clypeals 4 

 or 5. Palp to face 79-85. AR 1.6-1.8; A )3 750 um long. SVo sigmoid, 

 narrow, and smooth (Fig. 8). P 3 with ta 4 125 um long and LR 64. Body 



4.3-4.8 mm; wing 3.5 mm S. spinifera Tosio 



AnP longer with two unequal spines apically (Fig. 11). Clypeals 1 1. Palp 

 to face 89. AR 3.0; A 1095 um long. SVo band-like with fine spinulae 

 on surface (Fig. 12). P 3 with ta 4 195 um long and LR 58. Body 5.5 mm; 

 wing 4.3 mm S. macrocera Tosio 



PUPAE 



1. L-setae of VII-VIII subequal and not darker than those on preceding 

 segments and L 2 with 8 branches or less. D 4 of III-VII larger than D 5 

 and sometimes bifid on VII or VIII. Shagreen near apophyse of Til 

 weakly grouped. Thoracic spiracle (bulb) without spinulae. Coloration 

 yellow-brown with transparent-brown setae and spines, and muscle scars 



clear or colored (fulva-gp.) 2 



L-setae of VII-VIII shorter (by 1/3) and darker than those of VI and L 2 

 with more than 8 branches. D 4 of III-VII simple and subequal to D 5 . 

 Thoracic spiracle (bulb) with spinulae. Muscle scars dark (zavreli-gp) ... 3 



