50 Jan S. Doughman 



Leg segment lengths: 



















fe ti 



tal 



ta2 



ta3 



ta4 



ta5 



LR 



BV 



SV 



PI 1426 1674 



1472 



628 



426 



194 



163 



0.88 



3.2 



2.1 



P2 1542 1581 



876 



473 



310 



147 



155 



0.54 



3.7 



3.6 



P3 1814 2030 



1240 



643 



395 



170 



162 



0.52 



3.7 



3.1 



Tibial spur lengths: P^O, P 2 71 and 72, P 3 62 and 96. 

 Spiniform setae of ta )5 often paired: P^one, P 2 12-14,2-3,0,0,0, and 

 P 3 13-16,2-5,0,0,0. 



Tergite IX with each half with two groups of 6-8 moderately long setae; 

 anal point naked, awl-like, 70-90 long. Transverse sternapodeme thickly 

 rectangular. Straight coxite portion of phallapodeme, 70-170 long, 

 articulating with a strongly sclerotized, narrow, sigmoid SVo and a 

 membranous, ovate, setigerous MVo (Fig. 3). Gonocoxite, gonostylus 

 and other details typical for the genus, and found in the diagnostic de- 

 scription above. 



Material. — USA: California, Marin Co., Mill Valley. 12 Apr. 

 1957. Leg. H. L. Mathis, light trap. Holotype, U.S. National Museum 

 no. 65522 (poor mount). Oregon, Benton Co., Berry Creek (Willamette 

 R. basin), 9 mi N of Corvallis, 60 m alt. 17-24 Mar. 1960, leg. D. 

 Hansen; det. Hansen, 1973. 2 males. Univ. Minn. Collection nos. 

 DH69-280 and -281 (in balsam?). 



Remarks. — The holotype of diastena is very near fulva, but these 

 new specimens show diastena can be much larger and darker than fulva. 

 I doubt fulva will be found in the western Cordilleran. The numerous 

 dorsocentral setal rows and forward running prealars, and the unique 

 presence of setae on R 4+5 (Fig. 1), demonstrate that this species is the 

 most plesiomorphic species known in Sympotthastia. Its SVo is identi- 

 cal to spinifera and differs slightly from fulva. 



The presence of a species different fr om fulva in the western Cordil- 

 leran is also demonstrated by the pupa found in Waterton National 

 Park, Alberta, by Saether (1969) and keyed here as Sympotthastia sp. 

 Also, the larva from the Alaskan arctic, S. cf. fulva, is separable. It may 

 be that these unconnected metamorphic stages represent a single Cor- 

 dilleran species. 



Sympotthastia spinifera Serra-Tosio 

 S. spinifera Serra-Tosio, 1968:134-140, orig. design. (Figs. 1-4 of hy- 

 popygium; ecology); Serra-Tosio, 1971:224-227 (PL 123.2 phallapo- 

 deme; PL 127 hypopygium; Pis. 128-129 pupa; p. 277 ecology); Ferra- 

 rese and Rossaro, 1981:77-80 (larva description, Fig. 36 with mentum, 

 labrum, antenna, premandible, procercus; pupal description, Fig. 37). 



Sympotthastia macrocera Serra-Tosio 

 S. spinifera forma macrocera Serra-Tosio, 1968:137-138. 



