1 10 Gary S. Morgan 



trees provide the natural roosting site for many species of Neotropical 

 molossids, and it seems reasonable to hypothesize a similar roosting 

 ecology for T. brasiliensis in Florida prior to the extensive construction 

 of buildings. The probable tree-roosting habits of T. brasiliensis, coupled 

 with their extremely rapid flight, would limit predation and help to 

 explain their absence from most Florida fossil sites. 



Eumops glaucinus floridanus (G. M. Allen, 1932) 



Referred material— Stratum 3-V7222, partial edentulous left man- 

 dible; V7224, proximal end of right radius; V7226, proximal end of left 

 radius; V7227, one proximal and one distal end of femur; Stratum 

 unknown- V7220, right maxilla with P 4 -M 3 ; V7221, left mandible with 

 p3-ni3; V7223, proximal end of right humerus; V7225, proximal half of 

 left radius. 



Recent distribution. — Eumops glaucinus has the most restricted 

 distribution of any bat in the United States, being known only from 

 Charlotte and Dade counties in southernmost Florida. Wagner's mastiff 

 bat also occurs in tropical America from southern Mexico south 

 through Middle America, much of South America, and Cuba and 

 Jamaica in the Greater Antilles. The species has a disjunct distribution, 

 as it is not known to occur between southern Florida and southern 

 Mexico. 



Fossil record. — The fossil record of E. glaucinus is restricted to 

 Florida, where it is known from Vero, Monkey Jungle Hammock (Mar- 

 tin 1977), and the late Pleistocene Melbourne fauna, Brevard County 

 (Allen 1932; Ray et al. 1963). The fossil records from Brevard County 

 and Indian River County (this paper) extend the known range of the 

 species in Florida some 200 km north. 



Descriptions and comparisons. — The cranial and postcranial ele- 

 ments here referred to E. glaucinus are from a very large bat, and thus 

 need only be compared with the three largest species found in Florida — 

 E. glaucinus, Eptesicusfuscus, and Lasiurus intermedius. A maxilla and 

 partial rostrum agree with E. glaucinus and differ from E. fuscus and 

 L. intermedius as follows: presence of a tiny peg-like P 3 , stronger hypo- 

 cone on M 1 and M 2 , lack of a deeply incised nasal notch, vertical slit- 

 like infraorbital foramen, and vertical orientation of rostrum dorsal to 

 orbit, reflecting deep, laterally compressed snout (rostrum is dorsoven- 

 trally flattened in the two large vespertilionids). The mandible with p3- 

 m3, can be differentiated from E. fuscus and L. intermedius as follows: 

 presence of only two tiny incisors that are crowded between the canine 

 and mandibular symphysis, double-rooted p3, P3 and p4 subequal in size, 

 and the posterior margins of trigonid and talonid on molars not at right 

 angles to long axis of tooth row. 



