New Milliped Records 1 53 



On the gonopods (Figs. 2, 3), the subapical process of the secondary 

 tibiotarsus is reduced from the configuration depicted by Hoffman (Figs. 

 7c and 8c, pp. 201 and 203), so that it appears as a small tooth on the 

 main stem of the structure. It may be blunt or pointed (Figs. 4.-9) and is 

 absent from one male from Chester County (Fig. 9). Similarly, the apical 

 portion of the secondary tibiotarsus varies and may be blunt or pointed 

 and curved distad or ventrad (Figs. 4-9). The primary and secondary 

 tibiotarsi are subequal in length in specimens from North Carolina and 

 York, Chester, and Newberry counties, South Carolina, but the latter is 

 slightly longer in specimens from the southern part of the range. Den- 

 ticulations along the anterior edge of the primary tibiotarsus vary and are 

 essentially absent from some gonopods. The coxal apophysis and the 

 remainder of the gonopod are as described by Hoffman (1958). 



This increased knowledge of variation of P. c. incursus does not affect the 

 status of P. c. denticulatus Chamberlin, which occurs in the Piedmont 

 region of north-central Georgia. The two subspecies are distinguished by 

 several features, most notably the configuration of the coxal apophysis, 

 which is tri-lobed in P. c. incursus and smoothly rounded with the distal 

 margin entire in P. c. denticulatus. 



The range of P. crassicutis incursus can thus be expanded to include the 

 Kings Mountain region of North Carolina and the Piedmont and Coastal 

 Plain of southern South Carolina (Fig. 10). Efforts to find the diplopod in 

 North Carolina adjacent to the Kings Mountain region have been unsuc- 

 cessful, and this area therefore represents the known northeastern range 

 limit of the genus. 



ACKNOWLEDGMENTS.— We thank John C. Clamp and William W. 

 Thomson for assistance in collecting Pachydesmus from North Carolina; 

 Howard V. Weems, Jr., for access to material in the FSCA; and Norman 

 I. Platnick, for access to that in the AMNH. Specimens from Kings 

 Mountain, Barnwell, Baker Creek, and Hickory Knob State Parks, South 

 Carolina, were collected with permission of the S. C. Department of 

 Parks, Recreation, and Tourism, Division of State Parks. The photograph 

 is courtesy of Curtis Wooten, N. C. Wildlife Resources Commission. 



LITERATURE CITED 



Hoffman, Richard L. 1958. Revision of the milliped genus Pachydesmus 

 (Polydesmida: Xystodesmidae). Proc. U. S. Natl. Mus. 108: 181-218. 



Wray, David L. 1967. Insects of North Carolina, Third Supplement. N. C. Depart. 

 Agric. Div. Entomol., Raleigh 181 pp. 



Accepted 8 November 1978 



