154 Janice Heard Tardell, Richard C. Yates, David H. Schiller 



stage to shrub bogs. Apparently, H. andersoni populations thrive where 

 these two habitat components occur together. 



In this paper we report new information on the distribution and 

 habitat of H. andersoni in Chesterfield and Marlboro counties. During 

 the study we found the species inhabiting seepages in shrub and herb 

 bogs, a predictable habitat, but also found it in seeps of electrical 

 transmission line and gas pipeline rights-of-way, clearcut areas, and in a 

 broomsedge field. 



METHODS AND MATERIALS 

 Study Area 



We discovered Hyla andersoni in Marlboro County during a power 

 plant feasibility study for Carolina Power and Light Company. We then 

 investigated its occurrence along the drainages of Whites, Wolfs, Hicks, 

 Marks, and Phils creeks in northwestern Marlboro County and along 

 the Juniper Creek drainage in Chesterfield County. 



Most of the area investigated lies in the northern part of South 

 Carolina's sandhill region and is located east of the dividing line 

 between the Piedmont and Coastal Plain physiographic provinces. The 

 dominant vegetation of the area is the longleaf pine-turkey oak- 

 wiregrass (Pinus palustris-Quercus laevis- Aristida stricta) association. 

 Loblolly pine, P. taeda, becomes codominant with longleaf, especially in 

 moister areas. In the drier areas post oak, Quercus stellata; and black- 

 jack oak, Q. marilandica; become more numerous. In several large areas 

 south of Whites Creek the natural forests have been logged, cleared, and 

 replaced with monocultures of slash pine, P. elliottii, or left to regener- 

 ate naturally. Some of these areas were in the first or second season of 

 growth. Along the small stream basins that drain the sandhills, the vege- 

 tation is composed of "pocosin" or "evergreen shrub" species. The dom- 

 inants are sweet bay, Magnolia virginiana\ titi, Cyrilla racemiflora; var- 

 ious members of the heath family (Ericaceae); and other evergreen 

 shrubs, along with poison sumac, Rhus vernix. In many places the vege- 

 tation is covered with a tangle of greenbriers, Smilax spp. 



Methods 



United States Geological Survey 7.5 minute series topographic 

 maps were used to locate stream headwaters and intermittent streams. 

 These and other likely areas were inspected during the day to identify 

 appropriate habitat. We returned to the preidentified sites during the 

 evening to listen for the call of H. andersoni. Voice identifications were 

 verified by following the sounds to their source and observing calling 

 males, whose numbers were then estimated. For our purposes, a locality 

 where H. andersoni was found is one that is separated from another 

 locality by an area where the frogs do not call. The localities were 



