Hyla andersoni in South Carolina 155 



widely separated and generally were located along different tributary 

 streams or seepages. 



During 1978 we searched appropriate habitats from 11 through 22 

 July and from 25 through 27 July. Most effort at that time was directed 

 at the shrub bog habitat. We began our 1979 searches in May and con- 

 tinued them through the summer. During this time we discovered that 

 H. andersoni occurred in some clearcuts and transmission line rights-of- 

 way, so emphasis was placed on searching for and surveying these areas. 

 However, this did not preclude searches in the typical shrub bogs. All 

 Marlboro County localities found in 1978 were rechecked in 1979 to 

 investigate the continued presence of H. andersoni, but the Chesterfield 

 County locations were not revisited. 



RESULTS AND DISCUSSION 



Locations 



Hyla andersoni was first found in Marlboro County on 20 June 

 1978 in a shrub bog located in the headwaters of a tributary of Whites 

 Creek. In all, 67 localities were identified in Marlboro (56) and Chester- 

 field (11) counties during the summer of 1978 (41) and 1979 (26) (Figure 

 1). The continued presence in 1979 of H. andersoni was verified at 24 of 

 the 30 1978 localities in Marlboro County. At the six sites (all well- 

 developed shrub bogs) where H. andersoni was not found again in 1979, 

 the habitat was undisturbed. No human activities were evident. 



Habitat 



Plant species composition at most of the 67 sites was similar to that 

 reported by Means and Moler (1978). The dominant species are listed in 

 Table 1. 



Of the 26 localities found in 1979, 8 were located along transmis- 

 sion line or gas pipeline rights-of-way. These areas were similar in struc- 

 ture to the shrub bog-herb bog habitat depicted by Means and Moler. 

 The herb bogs developed where the actual clearing for the rights-of-way 

 crossed a wet area, and the shrub bogs developed to both sides where 

 canopy trees had been removed. Since the rights-of-way are periodically 

 maintained, they would not undergo succession to a closed canopy 

 forest that would preclude H. andersoni. 



Fifteen of the localities found in 1979 were in clearcuts to the east 

 of U.S. 1 and south of Whites Creek. These areas had been cut little 

 more than a year prior to our finding H. andersoni in them. We do not 

 know if the species inhabited these areas before the logging occurred, 

 but cursory checks in 1978 did not reveal their presence. The clearcuts 

 had herbaceous seepages surrounded by small shrubby stands at slightly 

 higher elevations; the shrub species were typical of older shrub bogs. 



