95 
glabrous coste and costule, by its scaly castes and in- 
dusiate sori. 
This distinct species is apparently rare and perhaps 
confined to the north-eastern provinces of Brazil where it 
was collected by SELLow (Campos Vittoria, no. 7332) and 
GoELp1 (RB, Parahyba) and once by Giaziou no. 15758 (B)- 
Dryopteris exculta (Mett.) C. Chr. comb. nov. 
Syn. Aspidium excultum Mett. Aspid. 69 no. 162 tab. 
17 fig. 9. 1858. 
Nephrodium eaxcultum Hook. sp. 4: 148. 1862, excl. 
syn. and most localities. 
Dryopteris effusa *exculta C. Chr. Ind. 263. 1905. Hieron. 
Hedwigia 46: 348. 
Aspidium laetum Moritz, Bot. Zeit. 1854: 855 (nomen, 
non Sw. 1817). 
_ Aspidium radicans Fée, Gen. 294. 1852 (ex descr., not 
Dryopteris Maxon 1908). 
Type from Venezuela, Tovar, in nemoribus, locis 
saxosis subapertis, Moritz no. 433 (B!). 
Closely allied to D. effusa, especially its var. divergens, 
which it resembles in most characters: rhizome, scales, 
division of lamina and kind of pubescence, but it can at 
once be distinguished from all forms of D. effusa by its 
sori being persistently indusiate; indusia rather large, 
red-brown, glabrous. Other but less reliable differences 
ra 20. D. acuta (Kl.) O, Ktze. 
Secondary pinnula from basal 
pinna, nat. size, and tertiary 
segment, 2: 
from D. effusa is the shape of the lamina, that is commonly ovate-elongated, long 
and gradually attenuated from a broad base towards the apex, not broadly deltoid 
as D. effusa. The rachis produces a scaly bud shortly below the apex. Lamina charta- 
ceous, brownish dark-green when dried (D. effusa var. divergens greyish green), up 
to 50 cm long, 30 cm broad below, generally tripinnatifid, in cutting very similar 
to D. effusa var. divergens, still the basal secondary pinnules are more broadly ovate; 
tertiary segments subacute. Underside, especially along the veins and between the 
sporangia, furnished with short, thick, cylindrical, yellowish hairs, which probably 
are glands with the juice dried out; rachises II are very suisse: pubescent beneath, 
mostly so towards their base (as in D. effusa typica). 
Venezuela: Tovar, Moritz nr. 433 (B, RB). 
Colombia: Fusagasuga, Striper nr. 503 (B); Popayan, LenMann s.n. (CC, RB). 
