oe ae 
var. guatemalensis (Bak.). 
Syn. Nephrodium guatemalense Bak. Syn. ed. Il. 498. 1874! 
Dryopteris guatemalensis Ktze. 1891; C. Chr. Ind. 269. 
Aspidium guatemalense Christ, Bull. L’Herb. Boiss. II. 6: 56. 1906 cum descr. 
Nephrodium xanthotrichium Sod. Rec. Cr. vase. quit. 52. 1883; Cr. vase. quit. 255! 
Differs mainly from typical D. exculta by its smaller size (lamina rarely 
more than 30cm long by 10—15 cm broad, often smaller), by its narrower secondary 
pinnules (scarcely 1 cm broad) and by its sharply acute tertiary lobes. Pubescence 
variable; most often the under surface is rather densely glandulose by thick, shining, 
red, glandular hairs that become pale-yellow when the juices dry out, and the 
rachises beneath are shortly pubescent by patent hairs, but in some specimens both 
glands and hairs are very few or totally absent. — N. guatemalense Bak. and N. 
xanthotrichium Sod. are to me quite identical. — Some specimens from Costa Rica— 
Panama come very near to typical D. exculta. 
Mexico: Chiapas, GuiespreGTH nr. 422 (Kew). 
Guatemala: Choelum (?), Satvin om GopMan 1862 (Kew, type of N. guatemalense Bak.); Alta Verapaz, 
Cubilquitz, v. TércKHEIM ed. D. S. nr. 8820 (RB, W — distributed as Nephrodium acutum 
Hook.); nr. II. 1088 (W), II. ob: (RB). 
Costa Rica: Llanuras de Santa Clara, Donnet SmitH nr. 6896, 6907 (W, both distributed as Nephro- 
um patulum Bak.); Pedra del Con gai PirTieR nr. 3548 (B,W):; El General, PITTIER nr. 
aa A, Los Palmares, PiTTieR nr. 10645 (W); Foréts de Tuis, Tonpuz nr. 11337 (W); Navarro, 
WercKELE (W); Juan Vifias, Reventazon cae Cook and Doy.e nr. 385 (W); Puerto Viejo, 
PiTTiER nr. 6928 (W, RB); Cerro Turubales, A. et C. BrapeE nr. 407 (R) 
Panama: Cana, WiL.iams nr. 880 (W); Porto Bello, Maxon nr. 5742? (W). 
Colombia: Bugava, Chiriqui, LeHMann s. n. (CC, RB). 
Ecuador: Chimborazo, Spruce nr, 5722 Kew); ad flumen Peripa, Sopiro (Kew, type of N. xantho- 
trichium Sod.). 
var. SQuUamifera n. var. 
Differs from all forms of D. effusa and D. exculta by its primary and secondary 
rachises being rather densely paleaceous by lanceolate, subentire, red-brown, thin 
scales; by its indusiate sori and glandulose underside it agrees with var. guale- 
matlensis. 
Costa Rica: Navarro, WerckLé nr. 16741. 16753, 16764 (RB). 
Obs. Fournrer-has referred some specimens from South Mexico to Aspidium 
Trianae Mett. (Mex. pl. I) which probably should be referred to D. exculta; a frag- 
ment of one of the quoted specimens (GuiEsBREGTH nr. 83 from Oxaca) belongs no 
doubt to this or a closely related species, very different from D. Trianae. It is larger 
than var. guatemalensis and without cylindrical hairs beneath. 
