254 ‘Mr. E. A. Newell Arber. Mew Species of [Jan. 10, 
axes, is readily intelligible on this view, and is unlike anything with which 
we are acquainted if these axes be really of the nature of shoots. Also, the — 
absence of any indication of foliar organs is remarkable on the latter 
supposition. Bearing in mind the fact that in the only known case in 
which we have any evidence as to the manner in which the seeds were 
borne in the Cycadofilices, that of JMedullosa, recently discovered by 
Mr. Kidston, they occur on the normal pinne of a frond, the foliar nature: 
of these branches in this instance is also highly probable. This conclusion 
is supported by the highly-compound nature of the frond attributed to 
Lyginodendron (Sphenopteris Heninghaust), from which, by reduction of the 
lamina, these branches might well be derived. In the case of Stur’s 
Calymmatotheca Stangert, a fossil recently discussed by Professor Oliver and. 
Dr. Scott,* the fructification is also borne on a frond. Lastly, it has been 
shown that the anatomical evidence of Lagenostoma Lomaxi “clearly indicates: 
that the cupule together with the pedicel was of a foliar nature.”"f We 
may, therefore, conclude with every degree of probability that the branched 
axes on which the seeds, Z. Sinclairi, were borne are of the nature of - 
segments of a compound frond with reduced lamina, and in view of the close 
affinity of this seed to that recently attributed to Lyginodendron, in 
addition to the characters of these branched axes themselves, this frond was. 
probably of the Sphenopteris type. 
Sununary. 
The following is a brief summary of the morphology of the seed, Lagenostoma 
Sinclairi, Kidston M.S. 
The seeds are radiospermic, and elliptical-oblong in shape. They vary from 
4 to 5°5 mm. in length, and 1°5 to 3 mm. in breadth. The testa is smooth, | 
and slightly ridged longitudinally. 
The integument at the apex is notched or fluted in a manner similar to the 
apical portions of Lagenostoma ovoides and L. Lomaxt. 
The seed was enclosed in a “cupule,” in many respects recalling that of 
L. Lomaxt. The length of the “cupule” is from 8 to 95 mm. The “ cupule,” 
at the stage of development represented in these specimens, entirely envelopes 
the seed. The “cupule” was a sac-like organ, prominently ridged longitudi- 
nally, and divided at the apex into a number of lanceolate, erect lobes. 
The seeds, enclosed in their “cupules,” were borne on the terminations of the 
finer branches of a highly compound frond with reduced lamina, in all 
probability of the Sphenopteris type. 
* Oliver and Scott, zbd., p. 230. 
+ Oliver and Scott, zbzd., p. 229. 
