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Current Herpetol. 20(2) 2001 



Table 4. Classifications of geoemydine 

 turtles of the family Bataguridae, derived from 

 results of this study. "*": species not examined; 

 "?": species of dubious generic allocation. 



Testudinoidea 

 Testudinidae 

 Emydidae 

 Bataguridae 

 Batagurinae 

 Geoemydinae 

 Geoemyda group 11 genera 34 species 



Genus Chelopus 2 species 



C. annulatus 

 C. rubidus 



Genus Cistocleminys 2 species 



C. flavomaiginata 

 C. galbinifrons 



Genus Ciiora 1 species 



C. amboinensis 

 C. aurocapitata 

 C. inccordi 

 C. pcini 

 C. trifasciata 

 C. yunnanensis* 

 C. zhoui 



Genus Cyclemys 4 species 



C. den lata 

 C. oldhamii 

 C. pulchristriata 

 C. tcheponensis 



Genus Geoemyda 4 species 



G. japonica 

 G. silvatica 



G. spengleri 

 ?G. leytensis* 



Genus Heosemys 3 species 



H. depressa* 

 H. grandis 

 H. spinosa 



Genus Leiicocephalon 1 species 



L. yuwonoi* 



Genus Melanochelys 2 species 



M. tricarinata* 

 M. trijuga 



Genus Notochelys 



N. platynota 

 Genus Pyxidea 



P. mouhotii 

 Genus Rhinoclenimys 



R. areolata 



R. diademata* 



R. fiinerea 



R. melanostenia* 



R. nasiita* 



R. pulcherrima 



R. punctiilaria 

 Maureinys group 

 Genus Maureinys 



M. annamensis 



M. caspicci 



M. iversoni* 



M. japonica 



M. leprosa 



M. mutica 



?M. pritchardi* 



M. riviilata 

 Genus Sacalia 



S. bealei 



S. pseudocellata* 

 S. quadriocellata 



1 species 



1 species 



7 species 



2 genera 1 1 



species 

 species 



3 species 



the Geoemydinae: 1 (0), presence of frontal 

 exposure on orbital rim; 12 (0), large and 

 elongated foramen palatinum posterius; 

 and 28 (2), axillary plastral buttress 

 connected around the portion between 

 peripherals and costals. Of these, however, 

 the first and the third were also observed in 

 various species of the Batagurinae (Table 

 1), Emydidae, and Testudinidae (Gaffney, 

 1979; Yasukawa and Hirayama, unpub- 

 lished data), and thus seemingly represent 

 primitive conditions of the Testudinoidea. 

 The second was observed in all geoemyd- 

 ines except for the Cistoclemmys-Pyxidea- 

 Geoemyda clade and thus may be a syn- 

 apomorph of the subfamily. However, we 

 failed to determine whether this state repre- 

 sents a derived or primitive condition, 



