YASUKAWA ET AL. — FllVl ()(iENY OF GEOEM\ DINT; TUR I i rS 129 



were elevated to lull species (i'liicliard, 

 1979; Prilehard and lichhau, 1984; 

 David, 1994). Like R. /'u/iciii/uriu and A'. 

 funerea, all ol ihc iliree species are 

 aquatic and have a sliallovvly notched 

 upper beak (Ernst and liarbour, I9(S9). in 

 addition, a small foramen on ihc \cniial 

 maxilla, which is suggested lo he a uniciue 

 synapomorphy of the A'. />iiiiciuiaria-R. 

 funerea clade (35 |l|), was recognized in 

 juvenile specimens ol R. iiielanosterna 

 and R. (liudcinuiu (Hirayama, unpub- 

 lished data). We thus leniativelv retain 

 ihc three species in ihc genus Rhinoclcninivs 

 (scnsii stricto). 



Direction of future studies on the 

 iieoeinydine pliyloiien v 



lii this siud\, we aitcinpicd as eomprc- 

 hensise an aiuiUsis as c\ci made lor the 

 phylogeiu ol ihc higliK dixcisiticd sublam- 

 ily Gcocmydinac b\ hcsi use o\ nioi jiholog- 

 ical inloi maiion . 



\n a icsuli, uc csiablishcd a besi- 

 liilmL' ph\ ioL'cncl ic lupoihcsis al ihis 

 level ll i!'s. ■? aiul 4), and acctirdingly 

 siii'i'csiod some chaiiL'cs iii ihe classifica- 

 lioii ol ihis siihlainiK (sec abo\c). Nev- 

 eilhclcss, oiii aiKiKsis lailcd U' iCNohe a 

 nunibci ol iiii'ioiip i clai loiislups. \iorc- 

 o\ei, oui lesulis, as well as our pre- 

 sum[")lioii ol ihc haui'.'unnc mouopluK, 

 sulisiaiii lalK coiiiiaJici ihc icnuIis of 

 leceni iirclinimai \ molcciilai ^^ullc^ on 

 ihc balaguiul plulogens (e.g.. Sues cl 

 al., 19S4: Shatter el al.. 199^; W u ci .d,, 

 l9i;S; McCoid CI ,d., :ui)li: 1 loiula cl ,d., 

 in press). In the luiuic, more compre- 

 hensive plulogenclic analyses of I'eslu- 

 dmoulca iism;' molecul.ii data are 

 desirable to lesi aiul rcMse, uhen neces- 

 sary, the lis jiothesis aiul elassilication 

 heiein piojiosed. 



A( k\( )\\ I i I H ,\1| Ms 



W'e wish lo e\piess i>ui gialitude lo 11. 

 \ \ndie\ss. 1. Das. R. Whilakci. K. 



Aoki, D. Aulh, \. Chigiia, R. 1. ( lomhie, 

 R. \V. Ileyer, A. II. \\\nn, (.. R. /ug, 

 II. l-.ndo, S.-I. Ueno, K. Ilaiooka anil Y. 

 Shibaia lor allowing us lo c\ammc speci- 

 mens in their cares, and lo R. Aoki, .\. 

 Araya, C. Blanc, 1. lla>aslu, S. Ikchara, 

 A. Kamata, S. Kaio, 7. Kuboya, isi. 

 Kame/aki, .A. Ko/.u, iVl. Malsui, A. 

 Mori, II. Ola, S. Scngoku, S. I anabe 

 and M. Tasumi lor providing precious 

 specimens. R. .Aoki, II. Oia, S. Scngoku, 

 M. Malsui and lasuiiii provided 



pertinent literal urc. We are also much 

 indebted to 1. Das, A. Mori, H. Ota and 

 M. Tasumi tor critical reading of the 

 manuscript. Ihis siud\ was partially 

 supported by a grant from I ujiuara 

 Natural History 1-oundaiion (to ^. 

 ^'asuka\\ a). 



Ll II R \ 1 1 Ki C'n 11) 



1{()MI\(.IR. Cl. A. 1889. Catalogue of 

 ilic ( iKUinians, Rh\ iichoccphalians, and 

 C rocodilcs in the Briiish Museum (Natural 

 History). Taylor and f-rancis, London. 



^1 I p. 



lioi I 1 Mil R. Cl. A. 1906. Descriptions of new 

 reptiles from Yunnan. .Ann. Mag. Naiur. 

 Hist. (7)17: 567-568. 



BoL K, R. 1980. Position systeinaiiquc dc 

 Geoclemys paliwannaiiiiinu Bourrcl. 1941 

 I Kcplilia-Testudincs-Emydidae). .Ainphibia- 

 Kcptilia 1(2): 14^) 159. 



Hoi Kki I. R. 1941. Los Toriues dc I'lndoch- 

 inc. Inst. Occanograph. I'lndochine 38: I- 



Hrwimii. I) \1 l'» * 1 mydid shell kinesis: 

 Mionicehanics and evolution. Copcia 1974: 

 707-727. 



BusACK. S. AND C. H. Ernst. 1980. Variation 

 in Mediterranean population of Maiirrmys 

 Gray. 1869 (Rcpiilin. Tcstudines. Emydidae). 

 Ann. Carncijic Mus \.itiir Hist 4«J 2<t- 

 264. 



C\RR. J. I . \NH J. \\ . BuKiivM. P^.st' l'ii>lo- 

 genetic iinplicalion» of karyotypic variation 



