Ciincnl //crpcloloi-y 20(2): 105-1.13, Dcccmlx-r 2(101 

 <<j 2001 by I he llcrpclological Society ol Japan 



Pliylogencfic Relationships of Cjeoeinydine I iirtles 

 (Kep(ilia: Bataguriclae) 



YuiciiiRou YASUKAWA,'* Ren HIRAYAMA,^ and Tsutomi- IIIKIDA' 



' IropiLul Hiosplu'tc Rcscaicli Cc/ilcr, Uiiivcrsily oj llw Ryiikyiis, Sishiliara, Okinawa 



903-02/3. JAPAN 



- Dcpariiiiciii o f In fonuaiioii. Teikyo Hcisei University, Uruic/o 22S9, Ichihara. Chiha 



290-0193, JAPAN 

 ^ Dc/H/n/iwni of /.oolo'^y, Graduate School of Science, Kyoto University, 

 Knushiriikawa-oiwukcclio, Sakyo-kii. Kyoto 606-8502, J. IPAN 



\l)slr;Kt: \l(>ii()|)h\ l\ ol llu' l):i(;imiri(l Mililainih ( .cooiin diruu' \\;is 

 iik'd. and lom-nclif ri'lalioiisliips uilhin llu' siihrainih \\i:rv iriti'irtd on llu- 

 hasis ol 35 morpholonkal iharailiTs. I wo approailus, parsimori> anal\sis 

 iisini; llic hraiu'li and honiid al^orilliin, and lU'i'^hhor-joininu ilnslerinu ol an 

 al)soluli' dislancc matrix, wcii' nsi'd. I hi- results <»!' these anal\ses \ielded 

 pli>lo^rams llial were almost identical in hraneliint; topoloji.N , and |)oorl> 

 supported llie iiioMopli\l\ ol ( leoeni) dinae. Mils Mil)l'aniil> thus seems to he a 

 metata\on. most likeh eonsisiiiiM ol (he sister uronp ol Mala^iirinae idco- 

 citiyild i^roiip) and a more primili\i' sloek ol |{alai;iiridae {\linircniy\ uroiip). 

 I he latter aeeoininodales \liiiircniy\ and Sacalia and ils inonopliNh is iiol \m II 

 supported. I he former consists ol the remaining ten i;eoem>dine genera uiiiiid 

 h> two s> napomorpliies ahseiuc or rednetion of the (|iiadraloju<;al . and pus- 

 leiiiuU short-sided anterior nenrals. Uelalioiiships re\ealed In our anal\ses 

 lurc'ssiiaU' sonie ehanues in the iieniiii elassil iealion of ( leoenndinae. I irsi of 

 all, ( i\i(ulcinin\ \ draN, I.S(i3 ilxpe species: ( /. Ihivunuir'^iiiiiiii. olleii •.\niiii\- 

 ini/i'd lo ( ttixii (.ra\, I.S55 M\pc speiics: ( (/. tti)\lu)iiicii\i\ \ . is shown in he a 

 \alnl m inis ilosi l\ n l.ileil In l'\\uli\i and ( nocniytld r.illu i Ih.in In ( linni 

 iM iisii sii u idi. I\lun(ii.lcnun \ \ 1 il/iniicr. IS.^S iiNpe species: A', piincliiliiriti) is 

 shinvii In he pi)l\ ph\ lelic, and ( hfl<)pu\ ( ope, 1H70 (t>pe species: ( . nihitlii^. 

 is ri-siirrecled In accn in mk mI:i le A', ntludii and A'. aiiitiilnUi. I he pl.i^li.il lim^e 

 seems in h;i\e evnUed lliuc linu's m llie Italauuridae in llu' ( ii/<v/m\- 

 \<>l(ic/u l\ cl.ide, ( iiiiiti iseiisii siiKini, .uul llic ( i\lnclctiliny \ l'\ \itltil clutle. 



Key words: Cicocniydinac; Hataguridac: Phylogcny; Ceoeinyda group: Mau- 

 remys group 



iMROni'mON 



* C oricspoiidmg .lutliot. IcI: » SI -9,S-iSy5- 

 8937; l ax: i 81 -9S'895'8%6. 



Imail address: h09O450((tjsci.u-ryukyu. ac.jp 

 (V, Vasukawa) 



.McDowell (1964) rccogni/cd ihrcx 

 sublaiuilies. Haingurinnc. Mmydinac. and 

 Tesfudininac. in the family Tc^ludmldac 

 Gaffiu-". iM,! \t.'.!.it) (loss* - us,-,' hose 



