Book I. PROCESS OF VEGETABLE NUTRITION. 171 



not exhausted at all. And hence also we may account for the phenomenon exhibited by plants vegetating 

 in distilled water under a receiver filled with atmospheric air, which having no proper soil to supply the 

 root with nourishment, effect the developement of their parts only at the expense of their own proper 

 substance ; the interior of the stem, or a portion of the root, or the lower leaves decaying and giving up 

 their extractive juices to the other parts. — Thus it appears that oxygene gas, or that constituent part of 

 the atmospheric air which has been found to be indispensable to the life of animals is also indispensable to 

 the life of vegetables. But although the presence and action of oxygen is absolutely necessary to the process 

 of vegetation, plants do not thrive so well in an atmosphere of pure oxygen, as in an atmosphere of pure or 

 common air. This was proved by an experiment of Saussure's, who having introduced some plants of 

 pisum sativum, that were but just issuing from the seed, into a receiver containing pure oxygene gas, 

 found that in the space of six days they had acquired only half the weight of such as were introduced at 

 the same time into a receiver containing common air. From whence it follows that oxygen, though the 

 principal agent in the process of vegetation is not yet the only agent necessary to the health and growth of 

 the plant, and that the proportion of the constituent parts of the atmospheric air is well adapted for the 

 purposes both of vegetable and animal life. 



758. Decomposition of water. Although the opinion was proved to be groundless, 

 by wluch •w ater had been supposed to be convertible into all the different ingredients en- 

 tering into the composition of the vegetable substance by means of the action of the vital 

 energy of the plant ; yet when water was ultimately proved to be a chemical compound, 

 it was by no means absurd to suppose that plants may possess the power of decomposing 

 part, at least, of what they absorb by the root, and thus acquire the hydrogen as well as a 

 portion of the oxygen which, by analysis, they are found to contain. This opinion was- 

 accordingly pretty generally adopted, but was not yet proved by any direct experiment. 

 Senebier pointed out several phenomena from which he thought it was to be inferred, but 

 particularly that of the gennination of some seeds moistened merely with water, and so 

 situated as to have no appai-ent contact with oxygen. The decomposition of water was 

 inferred also by Ingenhouz, from the amelioration of an atmosphere of common air into 

 which he had introduced some succulent plants vegetating in pure water. Saussure having 

 gathered a number of plants of the same species, as nearly alike as possible in all circum- 

 stances likely to be affected by the experiment, dried part of them to the temperature of 

 the atmosphere, and ascertained their weight ; the rest he made to vegetate in pure water, 

 and in an atmosphere of pure oxygen for a given period of time, at the end of which he 

 dried them as before, and ascertained their Aveight also, which it was thus only necessary 

 to compare with the weight of the former, in order to know whether the plants had in- 

 creased in solid vegetable substance or not. But after many experiments on a variety of 

 plants, the result always was, that plants when made to vegetate in pure water only, and 

 in an atmosphere of pure oxygen, or of common air deprived of its carbonic acid, scarcely 

 added any thing at all to their weight in a dried state ; or if they did, the quantity was too 

 small to be appreciated. But from a subsequent experiment, in which carbonic acid gas 

 was mixed with common air by the same experiment, the decomposition and fixation of 

 water by the vegetating plant is legitimately inferred. It does not appear, however, that 

 plants do in any case decompose water directly ; that is, by appropriating its hydi-ogen and 

 at the same time disengaging its oxygen in tiie form of gas, which is extricated only by 

 the decomposition of carbonic acid. 



759. Descent of the proper juice. When the sap has been duly elaborated in the leaf 

 by means "of the several processes that have just been described, it assumes the appel- 

 lation of the cambium, or proper juice of the plant. In tliis ultimate state of elaboration 

 it is found chiefly in the bark, or rather between the bark and wood, and may very often 

 be distinguished by a peculiar color, being sometimes white, as in the several species of 

 spurge, and sometimes yellow, as in celandine. It is said to be the principal seat of the 

 medical virtues of plants ; and was regarded by Malpighi as being to the plant what the 

 blood is to the animal body — the immediate principle of nourishment, and grand support 

 of life ; which opinions he endeavours to establish by the following analogies : if the blood 

 escapes from the vessels of the animal body, it forms neither flesh nor bone, but tumors ; 

 if the proper juices of the plant are extravasated, they form neither bark nor wood, but a 

 lump of gum, resin, or inspissated juice. The disruption of the blood-vessels and conse- 

 quent loss of blood, injures and often proves fatal to the animal. The extravasation of 

 the proper juice injures and often proves fatal to vegetables, unless the evil is prevented 

 by the skill and Uianagement of the gardener. Whatever m.ay be the value of these re- 

 marks as tending to establish the analogy in question, it cannot be doubted that the cam- 

 bium or proper juice constitutes at least the grand principle of vegetable organisation ; 

 generating and developing in succession the several organs of the plant, or furnishing the 

 vital principle with the immediate materials of assimilation. 



760. Tlie pi-oper juice is conveyed to the several parts of the plant hy an appropriate set of vessels. One 

 of the earliest and most satisfactory experiments on this subject, at least as far as regards the return of 

 the proper juice through the leaf and leaf-stalk, is that of Dr. Darwin, which was conducted as follows : 

 a stalk of the euphorbia heliscopia, furnished with its leaves and seed-vessels, was placed in a decoction 

 of madder-root, so as that the lower portion of the stem and two of the inferior leaves were immersed in 

 it. After remaining so for several days the color of the decoction was distinctly discerned passing along 

 the midrib of each leaf. On the upper side of the leaf many of the ramifications, going from the midrib 

 towards the circumference, were observed to be tinged with red ; but on the under side there was ob- 

 served a system of branching vessels, originated in the extremities of the leaf and carrying not a red but 

 a pale milky fluid, which, after uniting in two sets, one on each side the midrib; daaccndcd along with it 



