143 



the basal cell only by one sterile cell (fig. A — D). Sporangia silting immediately 

 on the basal cell I have not observed. 



The antheridia arise at the end of pecnliar narrow, cylindrical, colourless or 

 feebly coloured cells given olT from the apical end of the end-cells, not rarely from 

 cells sitting directly on the basal cell; in fig. P such a cell is even situated directly 

 on the basal cell. These androphore-cells, as they may be named, are not given 

 off in the same plane as the other branches but rise more or less vertically from 

 the horizontally directed cells. Usually one androphore only is given ofl' from the 

 same cell, but two androphores situated near each other also occur (fig. A'). As 

 mentioned below, the androjihore-cells may also be situated on the carpogonia. 

 The end of these androphore-cells gives rise to one or two antheridia. In the first 

 case a small cell, a little longer than broad, is cut ofT by a transverse wall (fig.F, G, A'), 

 in the latter the antheridial cells are cut off by inclined walls from the end of the 

 androphore-cell , leaving a little point between the two antheridia (fig. //, /); this 

 point may sometimes be lengthened into a short hair-like organ. 



As to the carpogonia and cystocarps, I am sorry to say that I have not arrived 

 at clearness, on account of the state of preservation of the material and perhaps 

 also because these organs occurred in very small number and in insufficient stages 

 of development. In particular it appeared difficult to find unquestionable tricho- 

 gynes. I think however that the cell shown to the right in fig. N is really a car- 

 pogonium with a spermatium attached to the trichogyne. In fig. P a cell-complex, 

 probably a young cystocarp, is seen bearing an androphore and quite near to it a 

 thin thread, which is perhaps a trichogyne, but no spermatium is attached to the 

 latter. A similar case is shown in fig. M, where a cell bears two thin, threadlike 

 organs, the one being certainly an androphore-cell, the other probably a tricho- 

 gyne. The great resemblance between the androphore-cells and the trichogynes 

 cause great difficulty, in particular when the antheridia are formed on the side of 

 the androphore-cell. Thus the case represented in fig. L might perhaps raise some 

 doubt. The resemblance between the filiform organ figured here and the andro- 

 phores represented in figs. H and / might perhaps suggest that it is an androphore 

 with two antheridia and prolonged point; the continuity of the protoplasmic con- 

 tents in the filiform organ and that of the cell from which it is given off' goes 

 however to prove, that these two organs belong together, being a carpogonium, 

 and that the two spermatia must have come from elsewhere and become attached 

 to the trichogyne. Small round cells looking like spermatia have sometimes been 

 found attached to various points on the surface of the plants (fig. A, 0, Q). In the 

 latter case (fig. Q) the small cell was adhering to a hyaline curved cell, the signi- 

 ficance of which I do not know. 



Of stages which could be supposed to be fertilized carpogonia or cystocarps I 

 have only found very few. The three-celled complex situated at the side of the 

 basal cell turned upwards in fig. P I regard as a young cystocarp. A similar 

 Ihree-celled stage is shown in fig. Q at the upper side, partly hidden by an over- 



