165 



outer small cells form an assimilatory tissue; the cells of the inner cortex are much 

 larger, containing also several bandlike and ramified chromatophores, but the total 

 mass of these bodies is small compared with the volume of the cell. Wille men- 

 tions these cells as store-cells, "Speicherzellen" (1887 p. 87), as floridean starch is stored 

 in great quantity in them. These cells are connected with each other by small pits. 



The cell-rows of the cortex depart from the longitudinal iilaments of the central 

 tissue, which consists not only of these filaments but also of irregular hyphse origi- 

 nating as outgrowths 

 from the barrel-shaped 

 cells of the inner cor- 

 tex (fig. 79). The dif- 

 ference between these 

 two kinds of iilaments 

 in the medulla has al- 

 ready been remarked 

 by older authors as 

 KuTziNG and Caspary; 

 Wille on the other 

 hand (1885 and 1887) 

 only refers to the se- 

 condary hyphse but not 

 to the primary longitu- 

 dinal filaments^ The 

 difference is conspi- 

 cuous, the longitudinal 

 filaments running very 

 regularly and consi- 

 sting of long cylin- 

 drical cells connected 

 with large pits, while 

 the hyphtic run irregularly, though chiefly in a transversal direction, and are com- 

 posed of more heterogenous cells, those of the proximal part being more or less 

 inflated, while the cells of the distal part are cylindrical. The cells of the hyphse 

 contain narrow, partly branched chromatophores. In the longitudinal filaments I 

 did not observe any chromatophores, but Denys (1. c. p. 10) states that their cells 

 contain colourless ones. This author states that the hyphaj are given off from the 

 longitudinal filaments'^; it is possible that they may also be produced by these, 



' For illustration of tlie anatomical structure of this species Wille gives only a cop3' of a figure 

 by Ki'iTziNG (Phj'col. gencr. tab. 72 fig. 6; Wille Taf. VIII fig. 14), representing Fiircellaria liimbrialis; 

 but this is identical with Pohjidcs rotundiis, which differs from Furcellaria just in the structure of the 

 medulla. Oltmanns makes the same error (1904 p. 546 fig. ;i30). 



^ Denys calls the longitudinal filaments "Langshyphen," but incorrectly, as these filaments have not 

 the active apical growth combined with slippinggrowthC'gleitendesWachstlium") characteristic of the hypha?. 



Fig. 79. 



Furcellaria fastlijiata. A, transverse sccliniv of frond, al llic liinil l)ct\vecn tlie me- 

 dulla and tlie inner cortex. B, longitudinal section of tlie same, c, inner cortical 

 celLs; Z, longitudinal filaments; h, liypIiSE. After living material, April. (190:1). 



