No. 486] 



PODOCARPINE.E 



359 



anticlinal divisions previous to being set free from the prothallial 

 complex, always retains its protoplasmic body as is generally the 

 case in other Gymnosperms, and thus cannot be confused with 

 any of the other contents of the microspore in tlie condition which 

 immediately precedes anthesis. 



In Podocarpus jerruginea from New Zt alaiid. iiiat<M-ial of which 

 we owe to the kindness of Dr. Cockayne, the carHer stages are 

 not so well represented as in the species described above, but so 

 far as they have been followed they present no essential deviation 

 from the course of events in P. polystachya. A figure 3, represents 

 the abietineous stage of development in this species. The preser- 

 vation is even less good than that of the Podocarpus already de- 

 scribed, and the protoplasm has shrunken from the microspore 

 membrane. In b figure 3, is shown a fully developed grain, in 

 which only one of the prothallial cells has undergone division. 

 The generative cell in this case is also free from divisions, although 

 it has rounded off and is almost ready to be set free from the 

 cavity of the microspore. C figure 3, presents a tangential view of 

 the first prothallial rudiment, which in this case has undergone 

 two anticlinal divisions, so that four cells have resulted. D figure 

 3, presents a longitudinal section through the air chambers and 

 shows anticlinal divisions in both of the prothallial cells. E figure 

 3, shows a similar condition" in the prothallial rudiments; but in 

 this case there are two lateral derivatives of the generative cell. 

 The latter are very small in size compared with the central cell of 

 the generative complex and with the derivatives of the prothallial 



As is represented in figures 3 and 4, starch is commonly found 

 in the pollen grains, especially in the younger stages, tliougli its 

 presence is by no means constant. A similar feature has been 

 noticed by Coker in the article cited above. 



n figure 4, shows the ganietophytic <levelo|.n.rnt in a probably 

 mature microsix.re of l)arr;/,!iuin Hi^hrlll!;, another n-proenta- 

 tive of the Fodocarpincie. The material in this cax- proved to be 



