No. 491] 



GYNAXDHOMORPHISM 



ain])le, a('('()r(liii<i to Boveri's view tlie siiiole inicleiis (that supplies 

 the male ]>art.s in the bee) is ilei-ived from the e^rg which in the 

 I)resent case contains tlie striped charac-ter; the other side is 

 derived from the conil)ined niu.4ei whicli slionld also the striped in 



contladictorv to the hxpothesls. On the other hand the facts are 

 wliat my Inpothesis ('alls for. 



Si) far 1 liave attempted to consider Toyama's cases witliout 

 regard to the (luestion of the sex of tlie right and left lialves because 



are more problematical, since we do not know in the moth the 

 nature of the factors that determine sex. Several {)ossibilities 

 must be considered. If however we are justified in extending the 

 conclusion reached above in regard to the origin of these gynan- 

 dromorph — a conclusion T repeat, that is reached independently 

 of the question of sex — to the case of the bee, where more is 

 known in regard to sex determination, we shall be led to some far 

 reaching and important considerations concerning sex determina- 



The moth that emerged from Toyama's gynandromorph cater- 

 pillar had ()n the left strii)ed half of the aixlomen, external female 



my view the right si(h' has come from tlie single spermatozoon. 

 It has prodticed the male sex. Two interi)retations are here 

 I)ossil)le. If tliere exist in the silk- worm moth two kinds of sperma- 

 tozoa—male and female producers — as shown by Stevens and 

 ^\'ilson for some other insects, the right side may be due to a 



