23 



Comments: Not mentioned in former lists (Williams & Francini, 1991; Cei 1993). Dixon (1983a :793) 

 mentions two individuals from El Bonito, Misiones and another one from Iguazii Falls in the materials and 

 methods section, but does not include Argentina in the distribution of the subspecies. Giraudo (1997) studied 

 21 specimens, all from Misiones. Velosa & Caramaschi (1993), in the 111 Latinoamerican Congress of 

 Herpetology, suggested that L. m. orinus (Griffm, 1916) and L. in. merremii (Wied, 1824) are synonymous 

 (then the Argentinian populations may be named L. merremii). As we do not know the results of the 

 publication of Velosa & Caramaschi, we use the taxonomy of Dixon (1983a). 



Liophis miliaris semiaiireus (Cope 1862) 



Opheomorphus merremi semiaiireus Cope, 1862. Proc. Acad. Nat. Sci. Philadelphia 14; 348. 



Liophis reginae ornata Jan, 1863. Arch. Zool. Anat. Fisiol. 2: 295. 



Opheomorphus fuscus Cope, 1885. Proc. Amer. Phil. Soc. 22: 190. 



Liophis miliaris semiaureus Dixon, 1983. Copeia 1983 (3): 792. 

 Type locality: Paraguay. 



Distribution: Buenos Aires, Corrientes, Chaco, Entre Rios, Formosa, southwestern Misiones and Santa Fe 

 (Giraudo, 1997; Giraudo & Quaini, 1997). Also Paraguay, Rio Grande do Sul in Brazil and Uruguay (Dixon, 

 1983 a, 1989; Achaval & Olmos, 1997). 



Comments: This taxon is probably a valid species (Giraudo, 1997). 



Liophis poecilogyrus (Wied 1825) 

 Coluber poecilogyrus Wied, 1825. Beitrage zur Naturgeschichte von Brazilien 1: 371. 

 Coluber m. nigrum Raddi, 1820. Atti. Soc. Ital. Sci. Modena 18: 38. 

 Natrix G.forsteri. Wagler, 1824. In Spix, Spec. Nov. Serp. Bras.: 16 

 Coluber doliatus Wied, 1825. Beitrage zur Naturgeschichte von Brazilien 1: 368 

 Liophis merremii var. sublineatus Cope, 1860. Proc. Acad. Nat. Sci. Philadelphia 1860: 252 

 Opheomorphus doliatus caesius Cope, 1862. Proc. Acad. Nat. Sci. Philadelphia 1862: 348. 

 Liophis omatissima Jan, 1863. Arch. Zool. Anat. Fisiol. 2 :53. 

 Liophis typhlus gastrosticta Jan, 1863. Arch. Zool. Anat. Fisiol. 2 : 53. 

 Liophis typhlus olivacea Jan, 1863. Arch. Zool. Anat. Fisiol. 2: 53. 

 Liophis verecundns Jan, 1863. Arch. Zool. Anat. Fisiol. 2: 53. 



Liophis cobella flaviventris Jan & Sordelli, 1866. Icon. Gen. Ophid. :18 pi. 5, fig. 92. 

 Liophis reginae var. viridicyanaea Jan & Sordelli, 1866. Icon. Gen. Ophid. : 18 pi. 2, fig. 91. 

 Rhadinaea dichroa Werner, 1899. Zool. Anz. 22 (581): 1 15. 

 Rhadinaea praeomata Werner, 1909. Ver. Vaterl. Naturk. Wurttenberg 65: 58. 

 Leimadophis poecilogyrus Amaral, 1927. Rev. Mus. Paulista 15: 78. 

 Leimadophis poecilogyrus reticulatus Parker, 1931. J. Linn. Soc. London 37: 285, pi. 16. 

 Leimadophis poecilogyrus platensis Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 77. 

 Leimadophis poecilogyrus xerophilus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 77. 

 Leimadophis poecilogyrus pictostriatus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 77. 

 Leimadophis poecilogyrus pinetincola Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 78. 

 Leimadophis poecilogyrus albadspersus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 78. 

 Leimadophis poecilogyrus montanus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 79. 

 Leimadophis poecilogyrus franciscanus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 80. 

 Leimadophis poecilogyrus amazonicus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 81. 

 Leimadophis poecilogyrus intermedins Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 81. 

 Leimadophis poecylogyrus lancinii Hoge, Romano & Cordeiro, 1976/77. Mem. Inst. 



Butantan 40/41: 75. 

 Dromicus poecilogyrus Lema, 1977. Iheringia (zool.) 50: 77. 



Leimadophis poecilogyrus Cunha & Nascimcnto, 1978. Publ. Avul Mus. Paraense E. Goeldi 31: 91. 

 Dromicus poecilogyrus Lema, 1980. Iheringia ser. zool. 55: 30. 



Liophis poecilogyrus Dixon, 1980. Milwaukee Public. Mus., Contr. Biol. & Geol. 31:13. 

 Type locality: Barra of Jucu, Espirito Santo River, Brazil. 



Distribution: Buenos Aires, Catamarca, Chaco, Cordoba, Corrientes, Entre Rios, Formosa, La Pampa, La 

 Rioja, Jujuy, Misiones, Salta, San Luis, Santiago del Estero, Santa Fe and Tucuman (Dixon, 1989; Dixon & 



