25 



Liophis reginae var. viridicyanaea Jan, 1866. Icon. Gen. Ophid. 18 pi. 2, fig. 1. 



Leimadophis poecilogyrus albadspersus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 78. 



Leimadophis poecilogyrus pinetincola Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 78. 



Leimadophis poecilogyrus amazonicus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 81. 



Leimadophis poecilogyrus franciscanus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 80. 



Leimadophis poecilogyrus intermedius Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 81. 



Leimadophis poecilogyrus montamis Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 79. 



Leimadophis poecilogyrus xerophilus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 77. 



Leimadophis poecilogyrus schotti Hoge, 1964. Mem. Inst. Butantan 30 (1960-62): 67. 



Liophis poecilogyrus Dixon, 1980. Milwaukee Public. Mus., Contr. Biol. & Geol. 31:13. (en parte). 



Leimadophis poecylogyrus lancinii Hoge, Romano & Cordeiro, 1916111 . Mem. Inst. Butantan 40/41: 

 75. [substitutive name to L. poecilogyrus amazonicus Amaral] 



Liophis poecilogyrus schotti Dixon & Markezich, 1992. Texas Jour. Sci. 44 (2): 153. 

 Type locality: South America, restricted to Sao Paulo by Hoge (1964), Mem. Inst. Butantan, 30, 1960-62 

 (1964): 68. 



Distribution: Northeastern Corrientes and Misiones. It is one of the most widespread subspecies, found from 

 Venezuela and Guyanas to the Amazonian basin, northeastern Brazil, southeastern Brazil except coastal 

 areas of Rio de Janeiro and Espiritu Santo (Dixon & Markezich, 1992). Paranaense province. 



Liophis poecilogyrus sublineatus Cope, 1 860 



Liophis merremii var. sublineatus Cope, 1860. Proc. Acad. Nat. Sci. Philadelphia 1860: 252 



Leimadophis poecilogyrus platensis Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 77. 



Leimadophis poecilogyrus pictostriatus Amaral, 1944. Pap. Av. Dep. Zool. S. Paulo 5: 77. 



Liophis poecilogyrus sublineatus Dixon & Markezich, 1992. Texas Jour. Sci. 44 (2): 157. 

 Type locality: Buenos Aires, Argentina. 



Distribution: Buenos Aires, Cordoba, southern Corrientes, Entre Ri'os, La Pampa, San Luis and central and 

 southern Santa Fe (Dixon & Markezich, 1992; Vuoto 1995). Also Rio Grande do Sul, Brazil and Uruguay 

 (Dixon & Markezich, 1992; Lema 1994). Pampeana and Espinal provinces (Dixon & Markezich, 1992; obs. 

 pers.). 



Liophis reginae macrosomus (Amaral 1935) 



Leimadophis reginae macrosoma Amaral, 1935. Mem. Inst. Butantan 9: 238. 



Leimadophis reginae maculicauda Hoge, 1952. Mem. Inst. Butantan 24 (2): 241. 



Liophis reginae macrostoma Dixon, 1983. Ann. Carnegie Mus. 52 (6): 1 13. 



Liophis reginae macrosomus Dixon, 1989. Smithsonian Herp. Inf. Serv. 79:21. 

 Type locality: Canna Brava, Goias, Brazil. 



Distribution: Northeastern Corrientes, Misiones and northern Salta (Dixon, 1983c, 1989; Alvarez et al., 

 1995; Giraudo, 1997). This subspecies is mentioned for boreal Chaco and for the Cerrado in Brazil, 

 Argentina, Paraguay and Bolivia. Distributed in South America east of the Andes from Colombia to 

 Argentina and Trinidad and Tobago (Dixon 1983 b). In Argentina it inhabits Paranaense and Yungas 

 provinces and transitional regions with Chaquena province. 



Comments: Dixon (1983 b: 137) mentioned that the two specimens from Salta are intergrades between L. 

 reginae semiliniata and L. reginae macrostoma. Williams & Francini (1991) included Formosa, probably 

 based on Dixon's maps (1983c, 1989) that show marks that apparently include Formosa, but the material 

 examined is from bordering localities of Paraguay. Alvarez et al. (1996) do not mention Formosa, and we 

 believe this record needs confirmation. 



Liophis sagittifer sagittifer (Jan 1863) 

 L.(iopeltis) sagittifer inn, 1863. Elenco Sist. Olid. Icon. Gen. Milano: 82. 

 Liophis pM/c/jer Steindachhner, 1867. Sber. Akad. Wiss. Berl. 55 (1): 267. 

 Rhadinaea sagittifcra Boulenger (/w/7////), 1894. Cat. Snakes Brit. Mus. Nat. Hist. 2: 165. 

 Zamenis argentinus Brethes, 1917. Physis 3 (13): 93. 



