﻿The 
  Anatomy 
  of 
  the 
  Female 
  Genital 
  Tract 
  of 
  the 
  Pupipara 
  etc. 
  27 
  

  

  It 
  was 
  v. 
  Siebold 
  who 
  first 
  deinonstrated 
  this 
  fact 
  (19). 
  Dufour 
  

   (4 
  and 
  5) 
  called 
  the 
  anterior 
  milk-glands 
  the 
  receptaculum 
  seminis; 
  

   Leuckakt 
  corrected 
  him. 
  

  

  But 
  it 
  is 
  not 
  the 
  entire 
  median 
  oviduct 
  which 
  is 
  thus 
  filled 
  with 
  

   sperm 
  and 
  functions 
  as 
  a 
  receptaculum 
  seminis, 
  but 
  the 
  dorsal 
  or 
  

   distal 
  end 
  of 
  it 
  (PI. 
  II, 
  Fig. 
  2 
  rec.sem) 
  which 
  is 
  considerably 
  wider 
  

   than 
  the 
  remainder 
  of 
  the 
  median 
  oviduct 
  and 
  forms 
  a 
  distinct 
  

   compartmeiit 
  in 
  it. 
  Leuckart 
  calls 
  this 
  the 
  Fundus. 
  Its 
  lumen 
  has 
  

   a 
  width 
  of 
  0,15 
  mm 
  in 
  the 
  adult 
  female 
  against 
  a 
  width 
  of 
  0,01 
  mm 
  

   in 
  the 
  portion 
  of 
  the 
  median 
  oviduct 
  immediately 
  adjoining 
  the 
  

   uterus. 
  It 
  is 
  slightly 
  arched 
  on 
  its 
  dorsal 
  surface 
  and 
  the 
  paired 
  

   oviducts 
  meet 
  it 
  to 
  the 
  right 
  and 
  left. 
  These 
  are 
  veiy 
  short 
  and 
  of 
  

   unequal 
  length. 
  When 
  in 
  either 
  ovary 
  an 
  egg 
  is 
  present 
  which 
  is 
  

   fully 
  matured 
  or 
  nearly 
  so, 
  as 
  is 
  the 
  case 
  with 
  the 
  left 
  hand 
  ovary 
  

   in 
  Fig. 
  11, 
  the 
  oviduct 
  on 
  that 
  side 
  is 
  apparently 
  almost 
  obliterated 
  

   through 
  the 
  crowding 
  of 
  the 
  egg 
  into 
  it 
  (PL 
  III, 
  Fig. 
  11): 
  when, 
  

   however, 
  the 
  largest 
  egg 
  is 
  still 
  small 
  and 
  immature 
  as 
  in 
  the 
  right 
  

   hand 
  ovary, 
  the 
  oviduct 
  on 
  that 
  side 
  may 
  have 
  a 
  length 
  of 
  0,23 
  mm. 
  

   And 
  between 
  these 
  extremes 
  we 
  find 
  all 
  intermediate 
  stages. 
  

  

  The 
  finer 
  structure 
  of 
  the 
  oviducts 
  presents 
  the 
  following 
  features. 
  

   The 
  structure 
  of 
  the 
  proximal 
  portion 
  of 
  the 
  median 
  oviduct 
  is 
  

   similar 
  to 
  that 
  of 
  the 
  uterus. 
  The 
  epithelium 
  of 
  the 
  fundus 
  and 
  of 
  

   the 
  paired 
  oviducts 
  is 
  peculiar 
  in 
  being 
  composed 
  of 
  very 
  high, 
  

   narrow 
  cells 
  giving 
  the 
  wall 
  a 
  much 
  greater 
  thickness 
  than 
  that 
  of 
  

   the 
  remainder 
  of 
  the 
  median 
  oviduct. 
  Surrounding 
  the 
  epithelium 
  is 
  

   a 
  network 
  of 
  muscle-fibres 
  similar 
  to 
  that 
  of 
  the 
  uterus 
  but 
  somewhat 
  

   thinner. 
  Their 
  arrangement 
  is 
  also 
  somewhat 
  more 
  regulär 
  than 
  that 
  

   of 
  the 
  uterus, 
  there 
  being 
  a 
  more 
  defmite 
  arrangement 
  into 
  circular 
  

   and 
  longitudinal 
  muscles. 
  The 
  ental 
  fibres 
  are 
  distinctly 
  circular 
  

   while 
  the 
  ectal 
  ones 
  are 
  longitudinal 
  although 
  their 
  arrangement 
  is 
  

   not 
  regulär 
  but 
  such 
  that 
  they 
  form 
  a 
  more 
  or 
  less 
  regulär 
  network 
  

   about 
  the 
  circular 
  fibres. 
  Among 
  the 
  muscles 
  are 
  numerous 
  connective- 
  

   tissue 
  fibres. 
  

  

  The 
  membrana 
  interna 
  is 
  very 
  delicate 
  in 
  the 
  median 
  oviduct. 
  

   In 
  the 
  paired 
  portions, 
  however, 
  it 
  is 
  very 
  thick 
  and 
  is 
  thrown 
  into 
  

   long 
  folds 
  which 
  are 
  longer 
  than 
  the 
  cells 
  they 
  abut 
  and 
  project 
  

   into 
  the 
  lumen 
  so 
  as 
  almost 
  to 
  fill 
  it, 
  leaving 
  only 
  a 
  very 
  narrow 
  

   passage 
  in 
  the 
  center 
  of 
  the 
  vessels 
  (PL 
  III, 
  Fig. 
  1 
  1 
  pa.ovi). 
  These 
  

   folds 
  are 
  all 
  inclined 
  toward 
  the 
  fundus 
  and 
  their 
  function 
  is 
  obvious: 
  

   they 
  form 
  a 
  System 
  of 
  valves 
  which 
  prevent 
  the 
  flow 
  of 
  the 
  sperma- 
  

  

  