16 



THE 



buds differ from the plants wliicli produce them. I have 

 described one of these examples in Bouvardia. The 

 pinkish white ''Bridesmaid" gives the red-flowered 

 ''Hogarth" from its root-cuttings. Three similar oc- 

 currences have been found in fancy Pelargoniums. The 

 root-cuttings of a white-flowered variety, "Pearl," give 

 a red-flowered form very like "Mme. Thibaut." "Mrs. 

 Gordon," which is a full rose-pink, with whitish edges, 

 gives from its root-cuttings flowers in which the two 

 posterior petals are marked with dark red, not unlike the 

 variety "Cardiff." A more striking case is that of 

 "Escot," which gives from its roots plants with bright 

 pinkish red marks, those of the original being purplish 

 red. The most curious feature of this case lies in the 

 increased size both of the plant and the flowers coming 

 from the roots, and it is scarcely possible to see the 

 petals of "Escot," which are characteristically rolled 

 back, side by side with those of the root-form, which are 

 not only larger, but also flat, without surmising that this 

 rolling back is an expression of the greater size of the 

 larger petal contained within the smaller, causing a want 

 of correlation between the growth of the inner and outer 

 tissues. 



Buckling or crumpling of leaves through want of cor- 

 relation was a conspicuous feature of some of Winkler's 

 "graft-hybrids," made from Solanum nigrum and 8. 

 lycopersicum, when the larger tomato was inclosed 

 within the smaller species. We have had a precisely 

 similar example in a salmon-fringed Pelargonium bred 

 by Mr. Jarman, of Chard. The leaves are obviously 

 buckled, the petals are laciniated, and the female parts 

 aborted, though the anthers are perfect. This male and 

 deformed flower is ].r()])tT to tlie outer tissues only; for 

 on two occasions tlic plants have produced shoots with 

 large flat leaves and normal liermaphrodite flowers with 

 their petals entire. Obviously, this normal plant was 

 inclosed within a skin of the fringed type. 



In all these examples, a somatic segregation has oc- 

 curred which determines the genetic potentialities. The 



