Is^o. 640] VARIATION AND HEREDITY IN LUPINUS 445 



allelomorph, so that dark flowers would then be linked 

 with light seeds. In either event plant VIII-27 would 

 be homozygous for dark flowers. It might have been 

 either homozygous or heterozygous for seed-coat color. 

 In the one case it would yield a progeny with dark-blue 

 flowers and light seed coats. In the other all plants 

 would have dark-blue flowers, but there would be three 

 light seed coats to one dark one. The results, however, 

 are both light-blue and dark-blue flowers. 



Pedigree VII is dark blue and has bred true for three 

 seasons. This spring culture VII-3-1 produced a single 

 white plant. It was the only plant from 77 seeds. The 

 parent plant was exceptional in that its seeds were lighter 

 in color than usual. 



Discussion. — It is realized that the facts presented in 

 the preceding pages are regrettably incomplete. It is 

 hoped, however, that they are sufticient to interest others 

 in lupines as suitable materials for i^enetic investigation. 



The striking pai'allelisni in tlK^ mutations occurring in 

 the two species, apriois and lunnis, is certainly a signifi- 

 cant })lionome]i()]i. All recent work with l)()tli plants and 

 animals iinncs that in xarictal ci'os^os homologous chro- 

 mosonu^s arc t'rocly int(M-('hanu-('alth' and that allelomor- 

 phic factors occupy identical h)ci in their respective 

 chromosomes. The work with multii)le allelomorphic 

 systems clearly indicates that a ])articular factor may 

 undergo a number of different changes. In L. apricus the 

 evidence at hand likewise indicates that the factors pro- 

 ducin- strijM'd wliite and liu'lit blue resi)ectively are eacti 

 alIelonioi-i>hi(^ to tliat for dark-i)lu(' Howcr color. Whether 

 they aiv allclonioiphic to one another remains to be 

 shown, thouuii that would he a piT.I)al)le supposition. 



From the data pivseiite,] in this ])aper it can not be 

 said whether tlower color and -eed coat color are both due 

 to the same factor or to linked factors. The latter is, 

 liowever, indicated })y two facts. In the first place we 

 already know more heritable ])atterns of coat color than 

 there are flower colors associated with them. At least 



