Contr. towards the Embryology and Anatomy of Polistes pallipes. 199 



some nuclei contain but a few large ones. In each Ducleus the 

 achromatin network is still seen and in these Strands are a number 

 of small, lightly staining, bodies similar in appearance to chromatin 

 granules. In the oldest Chamber many of the nurse-cells show a 

 layer of darkened cytoplasm lying around the nucleus (Fig. 63). This 

 we judge is due to the activity of the nurse cells. 



A little distal to the synapsis nuclei we find young oöcytes 

 similar to those we have already described. The nucleus has one, 

 sometimes two, large achromatin nucleoles, a cloudy and fibrillar 

 arrangement of other achromatin, in which lie imbedded a few 

 chromatin granules generally peripherally arranged (Fig. 64 a). The 

 large nucleolus becomes lost and does not appear again for some 

 time. The achromatin becomes arranged in threads (Fig. 64 ö), more 

 and more chromatin granules appear in these, and we finally find a 

 number of beaded chromosomes within the nucleus (Fig. 64 c). This 

 last stage is much scarcer than the others, no section ever showing 

 more than two and a majority of them none at all. Proximal to 

 these last nuclei lie those in synapsis. These show but a slight 

 degree of regularity and nowhere such a regulär polarity of the 

 Strands as Giardina (8 and 9) found in Dytisciis or Mantis^ the threads 

 in his figures also appearing more beaded. The greatest amount of 

 substance in the nucleus is arranged in a large irregulär mass in 

 which lie a number of distinct chromatin granules. Within this mass 

 could be Seen parts of the beaded threads, these would in part protrude 

 from it in all directions (Fig. 65). Just proximal to the synapsis 

 nuclei, we find the threads again becoming equally distributed 

 throughout the nucleus (Fig. 66). The nuclei are apt to become 

 elongated in form and will often lie two or three together (Paulcke) ; 

 later on, the shape changes and they separate from each other. From 

 here on for some little distance the threads retain their regularity, 

 and then break up, a process we will describe in the next stage. 



Passing down the tubule to the older oöcytes we find that the 

 oldest has increased very much in size, and both it, and the next 

 oldest, are elongated in the direction of the tubule. Besides the 

 nucleus, we now notice in each oöcyte a number of small bodies, 

 which, in their structure, are quite similiar to the nucleus (Blochmann^, 

 Stuhlmann, Korschelt, Paulcke, Gross). These have already 

 appeared in the third oldest oöcyte (Fig. 57), and here, in that which 

 is next to the oldest (Fig. 57, place 67), they are seen to lie in a 

 mass around the nucleus (Fig. 57) ; each has one or two small irregulär 



