No. 604] 



LINKAGE IN MAIZE 



235 



TABLE YIII 



Observed (from Table II) 125 



cu*r7in both 86^^°.^!".!..").^!^..°'!. 120.8 

 _curs onl/in the female5..°.^.!'!.°*!. 128.0 



Clearly, the first theoretical expectation fits the case 

 adequately, for with such a high value for P it is almost 

 certain that the deviations of the observed results from 

 the theoretical are due to errors of random sampling only. 

 Consequently one is justified in saying tliat crossing over 

 occurs in both male and female. This is especially true 

 when the fit in the second case is so poor. In fact, the 

 great difference between the two values for P makes it 

 seem reasonable that the intensity of the linkage is equal 

 in both male and female, although the high value for P 

 in the first case suggests that directly. 



Addition^al Linkages between Chlorophyll Factors 

 AND Aleurone 



Preliminary tests indicate that the same chlorophyll 

 factor, G, that is linked with R, is also concerned in a 

 linkage with one of the other chlorophyll factors termed 

 L. The latter has been found to be one of three factors, 

 two of which have already been described by Miles (1915), 

 involved in the production of chlorophyll in the seedling 

 stage of maize. 



Factors G and L seem to be linked although the data 

 from three back crosses, in which the numbers are small, 

 exhibit some variation in the percentage of crossing over. 

 Details of tliis linkage will appear in a later paper, deal- 

 ing witli tlic inlioritaiice of the three seedling chlorophyll 

 factors. 



Apparently, then, the factor pairs Rr, Gg, and LI con- 



