No. 605] 



.4.VI> CYTOPLASM 



291 



.si)erm enters the ef>'g abnormal inilolic limiics Mtid ali- 

 normal chromatin distribntioii arc direct l.\ con-clatcd 

 witli abnormalities in the larva\ all lioii-li the cu-- cylo- 

 plasra remains constant. Baltzer's line work ..n ^iicrl..< 

 crosses may also be cited as sIiowIiil:- lliat l.ddiV ;uid 

 (xodlewski's observations penetrated ()id\- part \va\ toward 

 tlie truth. 



As an example, only one of P.Mlt/.cr's need ))c 



cited. Wlien eggs of SplK, rrclni/iis were t'cililized by 



tril>idioii ()!■ cliromatin were normal niid the l.-irva' were 

 liild'iiK.riato. Hul tiiatcnial. I-'roin tlie rc.-iiiro.-al cross 

 ••iiiatrocliiio li>l.rl.U'' i nioM of tlum aluu.niinl ) did re- 

 sult, hill their inatcnial rcH'in!)laiicc was not due to cyto- 

 plasm. That it was due to iri-e-nlaritie> in tlie chromatin 

 distribution wa- proxed l.> r,Mll/.cr, followed the his- 

 tory of the maternal and paternal cliromosomes in tlie 

 hybrid embryos, lie found that the majority of the 

 paternal chromosomes (15 out of 18) were inactive at 

 the first cleavage. They were extruded from the develop- 

 ing oosperm nucleus, and degenerated. The cells of the 

 hybrid had then lM cin-oniosonies. IS nia{(>rnal am] pater- 

 nal, and their maternal roenihlaiice is e;i>lly explicable 

 on these grounds. It I< not explicahle on any other for 

 no abnorinal condition- obtained in eitlu-r c>toplasni or 



Ilcrh-t repeated the tlrM .>f r.:ilt/ei-^ cr-.-M- [SpJncr- 

 Prhmu.^y^Slnnnniioo i'f>'>tn^ ^) bnt .-lienilrMllx induced 

 parthenogenetic development in the liefoi-e tlie en- 

 trance of the si)erin. Tims tlie |.romicleu< w.i- l)ehind 

 at the first division and failed to ho inc.M-pornted into the 

 nucleus of one of tlie first daughter ceiU. ( )ne -ide of the 

 developing hybrid had, then, merely paternal ciiromo- 

 Manes, while the other had both matei-n.-d and i-atei-nal. 

 and in striking sequence to this distribution wwo enilir\ds 

 which had only ]iaterna] c1iara<-ler> on one >ide. wliiie 

 those of the otl'ier nde w.mv intermedinte. 



On the eviden.'e thu- far, Slmll him^df lia- not i-hu-ed 



