618 THE AMERICAN NATURALIST [Vol. LI 



numbers are not snfficiently large to form any conclusion 

 as to the intensity of the coupling, nor to establish the 

 existence of the same with certainty." 



A number of clearly segregating characters are known 

 in the tomato. Halsted lists 7 alternative unit character 

 pairs, while Price and Drinkard give 13. However, from 

 their own statements in regard to the behavior of these 

 characters, and from my own rather limited experience 

 with tomatoes, the number of different character pairs 

 which they list should be reduced. For instance, only two 

 allelomorphic pairs are known for color of fruit, viz., red 

 and yellow flesh or endocarp, and yellow and colorless 

 fruit skin or epicarp, while Price and Drinkard give four, 

 and Halsted three, character pairs of fruit color. Different 

 combinations of skin colors and flesh colors give the dif- 

 ferent colored fruits. For example, colorless epicarp over 

 red endocarp gives pink-colored fruit. 



TABLE VI 



Mendelian Characters in the Garden Tomato 

 (Revised from the lists <:i\on by Halsted and by Price and Drinkard.) 



It is somewhat uncertain as to the number of independ- 

 ent factors concerned in fruit shape. According to Crane 

 {loc. cif.) and Groth (1912 and 1915) there are a number 

 of factors and it is not always possible to distinguish be- 

 tween the various shapes. There is apparently a corre- 



3 See Crane, 1915, p. 4. 



