﻿Mr. 
  W. 
  Bateson. 
  

  

  may 
  perhaps 
  conjecture 
  that 
  this 
  element 
  has 
  been 
  transferred 
  to 
  the 
  male 
  

   side.* 
  

  

  The 
  presence 
  of 
  the 
  characteristically 
  masculine 
  comb 
  and 
  wattles 
  in 
  the 
  

   male 
  Sebright, 
  which 
  is 
  otherwise 
  henny, 
  shows 
  that 
  another 
  factor 
  similarly 
  

   detachable 
  governs 
  their 
  development. 
  

  

  To 
  the 
  breaking 
  up 
  of 
  large 
  compound 
  factors 
  the 
  production 
  of 
  inter- 
  

   mediate 
  types, 
  such 
  as 
  occur 
  among 
  the 
  colour-varieties 
  of 
  plants, 
  is 
  in 
  all 
  

   likelihood 
  due. 
  The 
  Sweet 
  Pea 
  and 
  the 
  Snapdragon 
  have 
  now 
  an 
  innumerable 
  

   series 
  of 
  such 
  colour-forms 
  which 
  may 
  be 
  represented 
  as 
  having 
  arisen 
  by 
  

   the 
  disintegration 
  of 
  the 
  various 
  anthocyanins. 
  That, 
  at 
  least, 
  is 
  the 
  simplest 
  

   way 
  in 
  which 
  their 
  origin 
  can 
  be 
  conceived. 
  

  

  To 
  the 
  final 
  result 
  many 
  qualifying 
  elements 
  contribute, 
  and 
  these 
  may 
  

   naturally 
  be 
  separate 
  entities. 
  But 
  change 
  in 
  the 
  amount 
  of 
  the 
  same 
  

   colouring 
  material, 
  and 
  diminution 
  in 
  the 
  mere 
  extent 
  of 
  its 
  distribution, 
  

   are 
  common 
  features 
  of 
  these 
  graduated 
  series. 
  As 
  the 
  cultural 
  develop- 
  

   ment 
  of 
  the 
  species 
  progresses, 
  more 
  and 
  more 
  of 
  these 
  quantitative 
  

   intermediates 
  appear, 
  and 
  are 
  selected, 
  until 
  a 
  practically 
  continuous 
  series 
  

   is 
  produced. 
  Although 
  the 
  interrelations 
  of 
  the 
  whole 
  series 
  can 
  be 
  

   represented 
  by 
  a 
  factorial 
  scheme, 
  the 
  assumption 
  that 
  each 
  of 
  these 
  factors 
  

   had 
  an 
  aboriginal 
  individuality 
  appears 
  gratuitous. 
  In 
  Antirrhinum, 
  for 
  

   instance, 
  the 
  ordinary 
  self-coloured 
  flower 
  segregates 
  as 
  a 
  single 
  unit 
  from 
  

   the 
  white. 
  But 
  there 
  are 
  " 
  Delila 
  " 
  forms 
  having 
  the 
  " 
  face 
  " 
  coloured 
  and 
  

   the 
  " 
  throat 
  " 
  white. 
  Another 
  variety 
  has 
  the 
  " 
  lip 
  " 
  coloured 
  and 
  the 
  

   peripheral 
  parts 
  white, 
  and 
  to 
  this 
  again 
  there 
  is 
  an 
  almost 
  exact 
  counter- 
  

   part 
  in 
  which 
  the 
  peripheral 
  areas 
  are 
  coloured 
  and 
  the 
  lip 
  nearly 
  white, 
  

   and 
  between 
  these 
  again 
  there 
  are 
  further 
  intergrades. 
  Apart 
  from 
  factors 
  

   modifying 
  its 
  quality, 
  the 
  colour 
  of 
  the 
  whole 
  corolla, 
  segregating 
  as 
  a 
  single 
  

   entity 
  from 
  the 
  white, 
  would 
  without 
  hesitation 
  be 
  represented 
  as 
  depending 
  

   on 
  a 
  single 
  factor. 
  Subsequent 
  experience 
  that 
  this 
  entity 
  can 
  break 
  up 
  

   into 
  an 
  indefinite 
  nnmber 
  of 
  fractions 
  is 
  not 
  evidence 
  that 
  the 
  original 
  

   representation 
  was 
  wrong. 
  This 
  reasoning 
  applies 
  to 
  a 
  great 
  range 
  of 
  

   phenomena. 
  

  

  In 
  view 
  of 
  the 
  chromosome 
  theory 
  of 
  linkage, 
  it 
  is 
  therefore 
  worth 
  

   remarking 
  that 
  we 
  do 
  not 
  find 
  linkage 
  especially 
  frequent 
  among 
  these 
  

  

  * 
  After 
  much 
  experiment 
  the 
  genetics 
  of 
  spur-development 
  are 
  still 
  very 
  obscure. 
  In 
  

   the 
  course 
  of 
  a 
  long 
  series 
  begun 
  by 
  crossing 
  spurred 
  Leghorn 
  hens 
  with 
  a 
  spurless 
  

   "Wyandotte 
  cock 
  (giving 
  ¥ 
  t 
  normal 
  in 
  both 
  sexes), 
  neither 
  form 
  has 
  reappeared 
  in 
  F 
  2 
  

   with 
  its 
  original 
  development. 
  Very 
  rarely 
  a 
  hen 
  with 
  minute 
  spurs 
  has 
  appeared, 
  and 
  

   occasionally 
  a 
  cock 
  with 
  the 
  spurs 
  sensibly 
  reduced. 
  Nor 
  from 
  spurred 
  hens 
  xF 
  1( 
  ^ 
  has 
  

   anything 
  approaching 
  the 
  original 
  types 
  been 
  raised. 
  Conceivably 
  the 
  detached 
  element 
  

   is 
  able 
  to 
  join 
  again 
  with 
  the 
  rest 
  of 
  the 
  female 
  complex. 
  

  

  