﻿366 
  

  

  Mr. 
  W. 
  Bateson. 
  

  

  periclinal 
  " 
  chimseras 
  " 
  of 
  white 
  over 
  green 
  which 
  produce 
  shoots 
  having 
  

   the 
  white 
  enclosed 
  in 
  the 
  green.* 
  To 
  these 
  must 
  now 
  be 
  added 
  the 
  cases 
  

   in 
  which 
  the 
  plants 
  arising 
  from 
  adventitious 
  buds 
  differ 
  from 
  the 
  plants 
  

   which 
  produce 
  them. 
  I 
  have 
  described 
  one 
  of 
  these 
  examples 
  in 
  Bouvardia. 
  

   The 
  pinkish 
  white 
  " 
  Bridesmaid 
  " 
  gives 
  the 
  red 
  flowered 
  " 
  Hogarth 
  " 
  from 
  

   its 
  root-cuttings. 
  Three 
  similar 
  occurrences 
  have 
  been 
  found 
  in 
  fancy 
  

   Pelargoniums. 
  The 
  root-cuttings 
  of 
  a 
  white 
  flowered 
  variety, 
  " 
  Pearl," 
  

   give 
  a 
  red-flowered 
  form, 
  very 
  like 
  " 
  Mine. 
  Thibaut." 
  " 
  Mrs. 
  Gordon," 
  

   which 
  is 
  a 
  full 
  rose-pink, 
  with 
  whitish 
  edges, 
  gives 
  from 
  its 
  root-cuttings 
  

   flowers 
  in 
  which 
  the 
  two 
  posterior 
  petals 
  are 
  marked 
  with 
  dark 
  red, 
  not 
  

   unlike 
  the 
  variety 
  " 
  Cardiff." 
  A 
  more 
  striking 
  case 
  is 
  that 
  of 
  " 
  Escot," 
  

   which 
  gives 
  from 
  its 
  roots 
  plants 
  with 
  bright 
  pinkish 
  red 
  marks, 
  those 
  of 
  

   the 
  original 
  beiug 
  purplish 
  red. 
  The 
  most 
  curious 
  feature 
  of 
  this 
  case 
  lies 
  

   in 
  the 
  increased 
  size 
  both 
  of 
  the 
  plant 
  and 
  the 
  flowers 
  coming 
  from 
  the 
  

   roots, 
  and 
  it 
  is 
  scarcely 
  possible 
  to 
  see 
  the 
  petals 
  of 
  " 
  Escot," 
  which 
  are 
  

   characteristically 
  rolled 
  back, 
  side 
  by 
  side 
  with 
  those 
  of 
  the 
  root-form, 
  

   which 
  are 
  not 
  only 
  larger 
  but 
  also 
  flat, 
  without 
  surmising 
  that 
  this 
  rolling- 
  

   back 
  is 
  an 
  expression 
  of 
  the 
  greater 
  size 
  of 
  the 
  larger 
  petal 
  contained 
  

   within 
  the 
  smaller, 
  causing 
  a 
  want 
  of 
  correlation 
  between 
  the 
  growth 
  of 
  the 
  

   inner 
  and 
  outer 
  tissues. 
  

  

  Buckling 
  or 
  crumpling 
  of 
  leaves 
  through 
  want 
  of 
  correlation 
  was 
  a 
  con- 
  

   spicuous 
  feature 
  of 
  some 
  of 
  Winkler's 
  " 
  graft-hybrids," 
  made 
  from 
  Solarium 
  

   nigrum 
  and 
  S. 
  lycopersicum, 
  when 
  the 
  larger 
  tomato 
  was 
  enclosed 
  within 
  

   the 
  smaller 
  species. 
  We 
  have 
  had 
  a 
  precisely 
  similar 
  example 
  in 
  a 
  salmon- 
  

   fringed 
  Pelargonium 
  bred 
  by 
  Mr. 
  Jarman, 
  of 
  Chard. 
  The 
  leaves 
  are 
  

   obviously 
  buckled, 
  the 
  petals 
  are 
  laciniated, 
  and 
  the 
  female 
  parts 
  aborted, 
  

   though 
  the 
  anthers 
  are 
  perfect. 
  This 
  male 
  and 
  deformed 
  flower 
  is 
  proper 
  

   to 
  the 
  outer 
  tissues 
  only 
  ; 
  for 
  on 
  two 
  occasions 
  the 
  plants 
  have 
  produced 
  

   shoots 
  with 
  large 
  flat 
  leaves 
  and 
  normal 
  hermaphrodite 
  flowers 
  with 
  their 
  

   petals 
  entire. 
  Obviously, 
  this 
  normal 
  plant 
  was 
  enclosed 
  within 
  a 
  skin 
  

   of 
  the 
  fringed 
  type. 
  

  

  In 
  all 
  these 
  examples, 
  a 
  somatic 
  segregation 
  has 
  occurred 
  which 
  determines 
  

   the 
  genetic 
  potentialities. 
  The 
  interpretation 
  that 
  they 
  are 
  periclinal 
  

   chimseras 
  is 
  probably 
  correct 
  for 
  the 
  most 
  part. 
  The 
  fringed 
  Pelargonium 
  

   is 
  obviously 
  of 
  this 
  nature. 
  Nevertheless, 
  the 
  fact 
  that 
  a 
  root-cutting 
  

   consistently 
  produces 
  a 
  certain 
  type 
  of 
  plant 
  which 
  is 
  not 
  the 
  original 
  

   does 
  not 
  prove 
  that 
  the 
  distribution 
  is 
  periclinal. 
  Another 
  possibility 
  is 
  

   well 
  illustrated 
  by 
  the 
  case 
  of 
  a 
  variegated 
  Spircea 
  ulmifolia, 
  having 
  the 
  

  

  * 
  ' 
  Jour. 
  Gen.,' 
  vol. 
  7, 
  p. 
  93 
  (1919). 
  

  

  