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  The 
  Genetics 
  of 
  Sex 
  in 
  Funaria 
  hygrometrica. 
  

  

  It 
  appears 
  evident, 
  therefore, 
  that 
  up 
  to 
  the 
  point 
  of 
  the 
  formation 
  of 
  the 
  

   female 
  organ, 
  the 
  cells 
  of 
  the 
  haploid 
  gametophytic 
  phase 
  retain 
  the 
  power 
  

   to 
  produce 
  monoecious 
  plants, 
  whereas 
  the 
  leaves 
  surrounding 
  the 
  male 
  

   organ 
  have 
  lost 
  this 
  power. 
  

  

  In 
  Funaria 
  in 
  the 
  early 
  condition 
  the 
  shoot 
  bearing 
  the 
  male 
  inflorescence 
  

   overtops 
  the 
  axis 
  bearing 
  the 
  archegonia, 
  and 
  this 
  appearance 
  leads 
  to 
  the 
  

   assumption 
  that 
  the 
  female 
  shoot 
  is 
  a 
  lateral 
  development. 
  From 
  specimens 
  

   taken 
  from^my 
  cultures 
  (see 
  text-figure) 
  the 
  point 
  of 
  oi'igin 
  of 
  the 
  female 
  

  

  Tracings 
  from 
  photographs 
  of 
  four 
  Funaria 
  plants, 
  showing 
  relation 
  of 
  female 
  shoot 
  to 
  male. 
  

   In 
  each 
  case 
  the 
  female 
  is 
  distinguished 
  by 
  the 
  protruding 
  "neck." 
  

  

  shoot 
  appears 
  variable, 
  and 
  in 
  some 
  cases 
  the 
  structure 
  simulates 
  a 
  dichotomy. 
  

   More 
  rapid 
  growth 
  of 
  the 
  male 
  axis 
  for 
  the 
  very 
  obvious 
  purpose 
  of 
  

   facilitating 
  fertilisation 
  would 
  lead 
  to 
  the 
  axis 
  bearing 
  the 
  archegonia 
  

   assuming 
  a 
  lateral 
  position. 
  In 
  view 
  of 
  the 
  new 
  observations 
  as 
  to 
  the 
  

   difference 
  in 
  potentialities 
  of 
  the 
  male 
  and 
  female 
  shoots, 
  it 
  becomes 
  

   important 
  to 
  determine 
  the 
  morphological 
  relation 
  of 
  the 
  two 
  parts 
  and 
  

   their 
  exact 
  cell-lineage. 
  For 
  at 
  some 
  critical 
  cell 
  division 
  a 
  change 
  takes 
  

   place 
  : 
  either 
  the 
  power 
  to 
  produce 
  the 
  monoecious 
  plants 
  is 
  extruded 
  in 
  

   the 
  female 
  shoot, 
  the 
  deficient 
  male 
  axis 
  continuing 
  the 
  development 
  ; 
  or 
  

   a 
  bud 
  which 
  has 
  lost 
  this 
  power 
  is 
  extruded 
  as 
  the 
  male 
  shoot, 
  while 
  the 
  

   female 
  shoot 
  retains 
  it 
  and 
  continues 
  the 
  development 
  of 
  the 
  axis. 
  

  

  In 
  this 
  connection 
  it 
  is 
  interesting 
  to 
  observe 
  that 
  Miss 
  E. 
  K. 
  Saunders 
  

   has 
  shown 
  me 
  some 
  of 
  her 
  preparations 
  of 
  the 
  termination 
  of 
  the 
  female 
  

   axis, 
  which 
  show 
  the 
  presence 
  of 
  antheridia 
  among 
  the 
  archegonia. 
  

  

  The 
  cytological 
  aspect 
  of 
  the 
  phenomena 
  has 
  not 
  been 
  studied, 
  and 
  the 
  

   generally 
  accepted 
  alternation 
  of 
  generation 
  within 
  the 
  group 
  with 
  its 
  n 
  

   and 
  2n 
  phases 
  has 
  been 
  assumed 
  throughout. 
  

  

  