494 Sir D. Bruce and others. Description of a 



the Commission that the strain was kept going in Pretoria in horses and 

 cattle, in which animals it produced the typical clinical symptoms and 

 post-mortem lesions associated with nagana, and that it was always regarded 

 as a pure uncomplicated strain of T. brucei. The thanks of the Commission 

 are due to Mr. Eobertson for his perseverance in sending inoculated animals 

 to their camp at Kasu. Like the Japanese general outside Port Arthur, as 

 one batch succumbed he sent on another, until at last he succeeded. 



The exact length of time this trypanosome was kept going at Pretoria 

 before being sent to Kasu is not given, but the information has been asked 

 for, and will be placed on record as soon as obtained. 



In the opinion of the Commission, the trypanosome dealt with in this 

 paper is the same as that discovered by Bruce in Zuhiland in 1894, and 

 named T. brucei by Plimmer and Bradford. Somkele is in the same district 

 in Zululand as that in which this species of trypanosome was first discovered. 



In this paper the old Zululand strain will be called the 1896 strain, that 

 being the year in which it was first described ; the new strain, the 1913 

 strain, the year in which it was received from Pretoria. 



The Zululand trypanosomes were described by Bruce in his original 

 paper* as heematozoa which vary among themselves a good deal in size and 

 shape. Photographs were also given, which show a distinct dimorphic type. 

 In a later paperf Bruce gives measurements of 200 trypanosomes taken 

 from preparations which had been made in Zululand in 1896, and also gives 

 six figures taken from the same source. From these it will be seen that 

 the trypanosome dealt with in Zululand in 1896 was a markedly dimorphic 

 form, with long and slender, intermediate, and short and stumpy forms. 

 Prom the above measurements and figures there cannot be the slightest 

 shadow of doubt about this. 



In 1896 Bruce sent this trypanosome to England, and it was at once 

 placed in the hands of Kanthack, Durham, and Blandford by the Eoyal 

 Society to be reported on. Their investigation lasted two years, and was 

 published in vol. 64 of the ' Proceedings ' of the Eoyal Society. In regard 

 to the shape of this trypanosome they state that " the Nagana parasites vary 

 considerably both in size and form ; they may be long and pointed or blunt- 

 ended and somewhat stouter ; some individuals are short and thick, with a 

 short flagellum, their protoplasm being crowded with rounded granules." 

 No one who reads Bruce's ' Progress Eeport ' and compares it with Kanthack, 

 Durham, and Blandford's 1898 report can doubt that the same trypanosome 

 was being dealt with. This trypanosome was distinctly dimorphic. 



* 'Further Eeport on the Tsetse-fly Disease, or Nagana, in Zululand,' 1896. 

 t ' Roy. Soc. Proc.,' B, vol. 83, p. 9 (1910). 



