Xo. 532] GENETIC AL STUDIES ON (EN OTHER A 229 



theorie." If this plant could be established as derived 

 from Oenotheras introduced into England by Messrs. 

 Carter and Co. at about 1860 from seeds said to come from 

 Texas, it would be a point of great importance, as will 

 appear in the following paragraphs. 



De Vries ( '05, pp. 384-385) offers strong evidence that 

 the strains of Lamarckiana at present cultivated in Eu- 

 rope have a genetic relation to seed of Messrs. Carter 

 and Co., of London about 1860. This seed is stated to 

 have been received unnamed from Texas and plants 

 grown from it were pronounced by Dr. Lindley to be 

 Lamarckiana. A specimen from one of these plants is 

 figured in * < The Floral Magazine, ' ' Vol. II, Plate 78, 1862, 

 this plate being reproduced in " L 'Illustration Horticole, ' ' 

 Vol. IX, Plate 318, 1862. This plate shows an (Enothera 

 with flowers about 10 cm. (4 inches) in diameter and 

 with a large amount of red coloration on the sepals and 

 ovaries; the stigma is figured both above and below the 

 tips of the anthers. The flowers of this illustration are 

 larger than those of Lamarckiana, as known to the 

 writer, and would do for grandiflora except for the posi- 

 tion of the stigma which is much closer to the anthers 

 than is typical for this species. The red coloration of 

 the sepals and ovaries is much too deep for typical La- 

 marckiana and not unlike some forms of grandiflora, but 

 the sepal tips, as drawn, are not so long or so pointed as 

 in the latter form. Indeed the identification of this plate 

 with any probable (Enothera is very difficult and the 

 reasons why it should be called Lamarckiana are to the 

 writer far from convincing, although it would perhaps 

 be as easy to argue for this identification as for any 

 other. 



It is, however, possible that new light may be thrown 

 on the composition of the cultures of Carter and Co. 

 through the plant in the Gray Herbarium described 

 above. The date of this specimen, 1862, together with 

 the very suggestive notes of Dr. Gray " from seed of 

 Thompson, Ipswich," and " said by English horticul- 



