No. 537] INHERITANCE OF THE "EYE" IN VIGNA 515 



end of the seed (see g, Fig. 1), is here called the Watson 

 Eye, from a variety known as Watson's Hybrid, which 

 has this type of eye. This variety, as well as the Hol- 

 stein variety, is said to have originated from crosses 

 between black varieties and Black Eye. My investiga- 

 tions show that both these color patterns occur in the 

 second generation of all crosses of this kind, as will be 

 seen later, thus confirming the supposed origin of both 

 these varieties. 



The Watson type of eye has not heretofore been recog- 

 nized. But since the essential difference between it and 

 other eye types was noticed, the writer has found in the 

 collection of cowpea varieties in the office of the agros- 

 tologist several races having this type of eye. The ex- 

 istence of this type was discovered as a result of the 

 discovery that in several crosses between eyed and com- 

 pletely pigmented races the ratio of fully pigmented to 

 individuals with eyes in F 2 was 3:1 in certain cases and 

 0:7 in others. The latter ratio suggested that two 

 genetically distinct types of eye were present in certain 

 crosses. A careful study of the rather extensive mater- 

 ial at hand revealed the types of eye already described, 

 and a tabulation of the statistical data relating to the 

 inheritance of each of them revealed the relations be- 

 tween these types which are discussed in this paper. 



Statistical Data 

 Cross: Small Eye X Solid Color (fully pigmented) 

 In all, 21 crosses of this kind were made. In all cases 

 Fj was solid color. It happened that in classifying the 

 individuals of the F 2 generation, before the types of 

 eye present were fully recognized, Large Eye and Small 

 Eye individuals were not separated in 5 of the 21 fam- 

 ilies. In the remaining 16 families, taken together, the 

 proportion of the various types of color pattern in F 2 

 was as follows: 



