No. 540] INHERITANCE OF COLOR IN CATTLE 715 



behave as dominant whites — i. e., they themselves are 

 white bnt sometimes throw red or black (not roan) calves. 

 The recessive white doubtless came in with the Dutch 

 flecked, the colored areas of which took the ' * differential 

 coloring" because they lacked the positive graying 

 factor; this recessive white must therefore be attributed 

 to a strain of partial albinism. The spotted color pattern 

 or coarse mosaic doubtless came in with the Dutch bulls 

 of the eighteenth-century importation. The areas com- 

 posing Group One are located about the two flank belts, 

 the underline, the median line and the face and a fine net- 

 work over the remainder of the body; those composing 

 Group Tivo cover the neck, sides, back, hind quarters and 

 legs in a network exclusive of the areas of Group One. 



w = Absence of such inhibitor. 

 R = Determiner for red pigmentation. 

 r = Absence of determiner for red pigmentation. 



With reference to Set No. 1, or group-unit No. 1, indi- 

 vidual cattle are gametically W 2 r 2 , WwR 2 or w 2 E 2 . With 

 reference to group-unit No. 2 they are w 2 R 2 , w 2 R r or 

 w 2 r 2 . There are therefore involving these characters 

 nine gametic and three somatic types of individuals, 

 which types are set forth in the following table: 



TABLE IV 



Nulliplex 

 Duplex 



Roan in this table stands for any animal with red and 

 white hairs interspersed, regardless of the proportion or 

 pattern. 



