402 



Mr. F. Galton's Experiments in Pangenesis. [Mar. 30, 



Litters subsequent to cross-circulation of both parents (common rabbits). 

 Average proportion of alienized blood in young a little less than j. 



kJXXii 01 



blood-mate 

 to 



-By 



blood- mate 

 to 



Number and character of litters. 



E 



R 



8 none Silver-grey, all like father or mother. 



E 





5 ditto. 



a 



o 



9 ditto. 



I* 



Q* 



8 ditto. 



J* 



Q* 



8 ditto. 







38 none Silver-greys. 



In another list (Table III.) I give particulars of all the litters I have 

 obtained from these rabbits, classified according to the operations which 

 the parents had previously undergone. 



I will now summarize the results. In the first instance I obtained five 

 does (A, B, C, D, and E) and three bucks (K, L, and M) which had un- 

 dergone the operation which I call w, and which had in consequence about 

 J of their blood alienized. I bred from these f, partly to see if I had 

 produced any effect by the little I had done, and chiefly to obtain a stock 

 of young rabbits which would be born with J of alien gemmules in 

 their veins, and which, when operated upon themselves, would produce 

 descendants having nearly | alienized blood (the exact proportion is 

 1 — (1 — J) 2 =-i-J). I obtained thirty young ones in six litters ; and they 

 were all true silver-greys, except, possibly, in one instance (out of the doe 

 A (u) by the buck M (w)), where one, of a litter of five, had a white fore leg, 

 the white extending to above the knee-joint. This white leg gave me great 

 hopes that Pangenesis would turn out to be true, though it might easily be 

 accounted for by other causes ; for my stock were sickly (both those on 

 which I had not operated and those on which I had suffering severely 

 from a skin disease), and it was natural under those circumstances of ill 

 health that more white than usual should appear in the young. 



Having, then, had experience in transfusion, and feeling myself capable 

 of managing a more complicated operation without confusion, I began the 

 series which I call w. I left my old lot of does untouched, but obtained 

 one new doe (G(w)), which had undergone the last operation, and three 

 bucks (K (u, w), M (u, w), N (u, w)) which had undergone both operations, 

 u and w. On endeavouring to breed from them, the result was unexpected, 

 they appeared to have become sterile. The bucks were as eager as pos- 

 sible for the does ; but the latter proving indifferent, I was unable to testify 

 to their union having taken place ; so I left them in pairs, in the same 

 hutch, for periods of three days at a time. Attempts were made in this 



* These rabbits belong to a breed liable to throw " Sandy " Himalayas, 

 f I always allowed the bucks to run for awhile with waste does before commencing 

 the breeding-experiments, that all old reproductive material might be got rid of. 



