No. 560] GENETIC AL STUDIES ON (ENOTHERA 459 



mained dwarfs, branching irregularly and presenting 

 varied forms of leaves ; there was also exhibited the same 

 wide range of flower size. The evidence was then clear 

 that in this family an F 2 plant of a type close to the mass 

 of the F 2 culture could throw off in the F 3 the same class 

 of dwarfs that appeared in the F 2 generation. 



The normal green rosettes constituting the mass of 

 the culture 12.52 (see Fig. 4, 12.52a) inclined strongly 

 towards the <i rand i flora parent of the cross, but presented 

 broader leaves not so strongly cut at the base. There 

 was a wide range of variation among the rosettes, and 

 forms appeared with narrow leaves which developed into 

 plants with a foliage markedly different from the mass 

 of the culture. The mass of the culture presented the 

 same evidence of progressive evolution which was shown 

 in the F 2 generation, i. e., the leaves were large and 

 crinkled as in the F 2 parent plant 11.41c, and there was 

 likewise maintained the same advance in the size of the 

 flowers, which ranged from types as large as or larger 

 than the grandiflora parent to types as small as that of 

 the F 1 hybrid lO.SOLa. In short, this F 3 generation, cul- 

 ture 12.52, from a plant 11.41c, fairly representative of 

 the mass of the F 2 , repeated the performance of the F 2 

 in exhibiting a large class of the same type of dwarf from 

 etiolated rosettes and also repeated very much the same 

 range of variation in leaf and flower characters shown by 

 the F 2 generation. There was then presented a behavior 

 closely parallel to that of Lamarckiana when it throws off 

 in successive generations the same marked types of vari- 

 ants, which hold true. 



Late in the season two plants were noted (12.52/a and 

 12.52/fr) upon which a large number of flowers were 5- 

 merous, t. e., the flowers had 5 sepals, 5 petals, 10 stamens, 

 and as far as noted 5 cells in the ovary. I am not aware 

 that this character has before been noted in the genus 

 CEnothera, These flowers were not restricted to partic- 

 ular branches and were found in the same inflorescence 

 with normal flowers. The 5-merous flowers, were not ob- 



