No.561] GEXETICAL STUDIES OX JENOTHEBA 553 



would be 1 : 1, provided that the female gametes R and r 

 were formed in equal numbers and mated in equal pro- 

 portions with the male gametes (r). It is at least clear 

 from this culture that the factor for red glands (R) is in 

 this case carried by a certain proportion of the female 

 gametes and that the female gametophytes for the plant 

 11.35m must be of two sorts (R and r). 



The plants of this culture, 12.45 (11.35 mXa), failed 

 to exhibit consistently the other differences associated 

 with the presence or absence of red glands as illustrated 

 by the two F x types 11.35m and 11.35a. There was a 

 marked progressive advance over the parent species, 

 biennis and grandiflora, in leaf size and general vigor, 

 but not in flower size, the petals ranging from 1.5 to 3.2 

 cm. in length. 



From the cross 11.35 a X m the contents of one capsule. 

 223 seeds, were sown. The culture. 1l'.44, gave 142 plants 

 which were brought to maturity. Of these 23 plants pre- 

 sented the red-colored papillae characteristic of 11.35m 

 and of the biennis parent, and 119 had the coloration of 

 11.35a and the grandiflora parent. The proportions of 

 these two types (23:119) is far from the expected ratio 

 1 : 1 on the hypothesis considered above, but it should be 

 noted that the total number of plants in the culture (142) 

 is small. The main consideration is, however, clear, viz., 

 that the factor for the red papillae is in this case carried 

 by a certain proportion of the male gametes and that the 

 male gametophytes from the plant 11.35m must be of 

 two sorts (R and r). Thus in both crosses (11.35 m X a 

 and 11.35 aXm) the character of the red papillae is repre- 

 sented in certain of the gametes both male and female and 

 the character is not sex-limited. 



The plants of the culture 12.44 (11.35 a X m) also failed 

 to show consistently the other differences associated with 

 the presence or absence in the F, of red papillae. Th 

 was a similar progressive advance over the parent spc 

 in leaf size and vigor, and likewise no advance in flo 

 size, the petals ranging from 1.3-3 cm. in length. 



