No. 561] GENETIC AL STUDIES ON OENOTHERA 



tween these extremes was an assemblage of intermedi- 

 ates impossible of classification. In short, this portion of 

 the culture presented an excellent illustration of a relative 

 segregation of characters, with the extremes, however, 

 quite far from the pure parent types. As the culture 

 grew to maturity a reclassification of the types became 

 necessary and finally five groups were separated as de- 

 scribed below. 



Group A consisted of 132 plants which had the La- 

 in a rcl'i ana -like foliage and short internodes (Fig. 16) 

 of the parent F, hybrid 11.35La, together with the 4- 

 angled buds and flower form of this plant. These plants 

 developed from the group of rosettes with broad crin- 

 kled leaves of the LamarcUana type. The size at matur- 

 ity ranged from plants 1.3 m. high to dwarfs 4 dm. in 

 height ; the habit and leaf size exhibited great variation. 

 The extreme types of dwarfs (13 in number) had very 

 much the habit of nanella. The flowers varied greatly 

 in size, petals 3.5-1 cm. long, with the stigma both above 

 and below the level of the anthers. There was. there- 

 fore, in this group a decided segregation of flower size. 

 A peculiar feature of these flowers was the very com- 

 mon cutting of the petals at the edge into narrow seg- 

 ments as in laciniate varieties of flowers. This is, as far 

 as I know, a new character in the genus (Enothera. The 

 greatest development of leaf size and extent of crinkling 

 observed in this group is illustrated in Fig. 17, which 

 shows two rosette leaves of one of the hybrids, 12.41Lp, 

 compared with the rosette leaves of the parent types of 

 biennis and grandiflora. 



Group B contained 5 dwarfs, 3-4 dm. high, sparsely 

 branched or not at all, and with narrow leaves. The 

 buds and flowers were grand ifi ora-Yike in form, but the 

 petals were only about 1.8 cm. long. These dwarfs were 

 very delicate and presented the characters of (Enothera 

 reduced in size to about the simplest terms. They re- 

 called the class of dwarfs in the F 2 from the plant 



