No. 547] GENETIC AL STUDIES ON (ENOTHEBA 



I have recently had the opportunity of examining two 

 herbarium sheets of American wild plants, referred to 

 in my earlier paper (Davis, '11, p. 227), which were 

 thought by De Vries ('05, p. 386) to be 0. Lamarckiana. 

 The first of these, at the New York Botanical Garden, is 

 a specimen collected by A. W. Chapman in Florida (1860 

 or earlier). Duplicates of this material are said to be at 

 the Biltmore Herbarium and at the Missouri Botanical 

 Garden (MacDougal, '05, p. 6). The second sheet, at the 

 Philadelphia Academy of Natural Sciences with an appar- 

 ent duplicate at the New York Botanical Garden, is of a 

 specimen collected by C. W. Short near Lexington, Ky. 

 This specimen was later considered by Miss Vail to be 

 0. grandiflora and a possible escape from cultivation. 

 There is nothing on these sheets from Florida and Ken- 

 tucky that is not represented in a fair range of herbarium 

 material of grandiflora such as may now be found in my 

 own collections and at the New York Botanical Garden. 

 In no point do the sheets closely approach Lamarckiana 

 except that they have large flowers. In justice to Pro- 

 fessor De Vries it should be stated that he expressed his 

 opinion before the rediscovery of the habitat of (Enothera 

 grandiflora by Tracy in August, 1904, and consequently 

 before there was available the extensive material of this 

 species now assembled. Could he have made the com- 

 parisons at present possible he would not, I am sure, 

 have given the opinion quoted above. 



There has then so far been found in the American 

 herbaria and records, and these have been very thor- 

 oughly examined by various workers, no evidence that 

 (Enothera Lamarckiana is at present or ever has been a 

 component of the American flora as a wild native species 

 There are in the south and west certain large-flowered 

 species of (Enothera of considerable interest because of 

 their possible affinities with grandiflora, and these should 

 be studied by those in a position to do so. A number of 

 these are represented in American herbaria; others m 

 the British Museum are referred to by Gates (>lla T p. 



