406 



THE AMERICAN NATURALIST 



[Vol. XL VI 



green (without red-colored glands). The flowers were 

 indistinguishable from those of the small-flowered types 

 of Lamarckiana except for the absence on the ovaries of 

 red-colored glands. They differed from the large-flow- 

 ered types (petals 4-4.5 cm. long) not only in size, but 

 also in the shorter stigma lobes which were not above the 

 tips of the anthers. My experience has shown that the 

 flowers in a second generation of these (Enothera hybrids 

 will present a wide range in their measurements, many 

 flowers surpassing in size those of the grandifiora parent 

 and equalling the largest forms of Lamarckiana. Conse- 

 quently there is good reason to be hopeful of obtaining 

 large-flowered types in the F 2 generation. 



6. Capsules. — The capsules, about 2 cm. long, were 

 stout and similar to those of Lamarckiana. The bracts 

 exhibited the deciduous tendencies of Class II, a 

 Lam arc k la n a - \ ike h abit . 



7. Seeds. — The seeds were of a shade of color inter- 

 mediate between the light and dark brown of the parents. 



Speculation as to the probabilities of the behavior of 

 these F 1 hybrids (11.35La, 11.35a, and 11.35m), and the 

 reciprocal crosses (11.35 a X m, and 11.35 m X a) in the 

 second generation, is not called for, since we hope in a few 

 months to have data on the questions involved. Some of 

 these problems concern the general behavior of species 

 crosses and are to the writer as interesting and im- 

 portant as the attempt to synthesize a Lamar ckiana-\\ke 

 hybrid between biennis and grandifiora. It may, how- 

 ever, be pointed out that there are no important char- 

 acters of Lamarckiana which are not represented in some 

 one of these hybrids or in their parents, except those of 

 the somewhat larger size of certain organs. Since my 

 cultures have shown that hybrids of (Enothera in the F 2 

 generation frequently present marked advances over 

 their parents in the measurements of petals, leaves, etc.. 

 these differences of size in an F x hybrid are likely to 

 prove more apparent than real. 



2. grandifiora Dx biennis D (11.32).— As would be 



