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THE AMERICAN NATURALIST [Vol. XLVI 



much variation from this plant in tlie measurement- 

 and proportions of the organs with the greater tendency 

 towards the grandiflora parent type. There was, how- 

 ever, no constancy apparent and plants with flowers 

 similar to grandiflora might present a foliage of a dif- 

 ferent type, most frequently in the form of larger and 

 strongly crinkled leaves. A few plants were so close to 

 grandiflora that taxonomically they would probably be 

 included in the range of this species, although they could 

 not be considered identical with the strain grandiflora B; 

 there were no segregates so close to biennis A. 



A count was made with the assistance of Mr. H. H. 

 Bartlett of the plants with green stems (biennis-like) and 

 of those with red stems (grandiflura-Yike). This was ex- 

 tremely difficult for the reason that the mass of the cul- 

 ture presented intermediate or mottled conditions similar 

 to the F t hybrid plant 10.30La. However, of the 1,310 

 plants, 195 were classified as red-stemmed and 192 as 

 green- stemmed; the red were markedly nearer the more 

 graudi flora-like end of the culture and the green nearer 

 the biennis end. The expected number according to 

 simplest Mendelian ratio should have been about 327 

 plants of each color. Considering that anthocyan is so 

 variable in its appearance, and consequently most un- 

 satisfactory for a study of color inheritance, these results 

 are by no means against Mendelian expectations. No 

 two persons independently would make the same count in 

 sueh a culture, for the results all depend upon what con- 

 ditions of the plant are denned as intermediate or 

 mottled. 



The culture presented a number of remarkable types 

 which no taxonomist would think of identifying with 

 either biennis, grandiflora, or the F, hybrid plant 10.30La. 

 These will not at present be described in their many 

 characteristics, but will be briefly listed. 



11.41c, a type with conspicuously crinkled leaves, repre- 

 sented by about 170 plants, apparently intergrading with 



