color character and the other with respect to the texture 

 of the endosperm. A self-pollinated second ear from 

 the plant that produced ear 471 had 24 horny seeds and 

 5 waxy, all of them white. A self-pollinated ear was 

 also secured from the plant which was the male parent 

 of both 471 and 472. This ear had 31 white seeds and 

 128 colored, all of them waxy. The total percentage of 

 white seeds in ear No. 471 was 48, a close approximation 

 to the expected 50. 



The nature of this experiment seems to preclude the 

 application of the theory of gametic coupling which 

 would explain correlations by assuming attractions or 

 repulsions between the character determinants in the 

 gametes. While attractions which might cause some 

 combinations to be represented by larger numbers of 

 gametes would disturb the Mendelian ratios, they could 

 not produce the effect of correlation or coherence of 

 characters. 



Thus the female parent might produce more gametes 

 bearing white and waxy than it did colored and waxy, or 

 there might be a greater fatality among the colored 

 waxy gametes. Any such disparity in the classes of 

 the gametes would result in more than 50 per cent, of 



