possibility of selective pollination did not prevent the 

 appearance of significant correlations. It appears no 

 more reasonable to assume that the number of individuals 

 that are to show a particular combination of characters 

 is always determined at the time the gametes are formed, 

 than it would be to argue that the number of nodes des- 

 tined to bear a juvenile type of foliage are definitely de- 

 termined at this time. 



The theory of gametic coupling advanced by Bateson 

 and Punnett leads them (1911, p. 6) to entertain the idea 

 that, while segregation is definite and complete, the ap- 

 parently significant cytological processes of maturation 

 may have nothing to do with the phenomenon. These 

 authors would have the association of characters deter- 

 mined before the maturation divisions. 



(127 + 1 + 1 + 127) it is nmst difficult to conceive 

 tern can be produced in the maturation-divisions of 1 



ub-epi.lorinal 



The suggestion that segregation occurs early in the 



ontogeny of the individual was apparently 



oned by 



the belief in the definiteness of the mathematical rela- 

 tions between different character pairs. As we have 

 seen, there is little evidence for this belief. The correla- 

 tion shown in ear 471, Table III, affords a definite indica- 

 tion that such associations of characters may be formed 

 after the production of the gametes. 



Even though the definite and material segregation ot 

 characters be maintained, all grades of correlation could 

 still be determined during the two nuclear divisions that 

 follow synapsis. The process might be looked at as 

 follows : 



When two character pairs are involved in a hybrid, the 

 tetrads resulting from the different mother cells would 



